1. Introduction
The National Museum of Costa Rica (MNCR) houses a patrimonial collection of Hymenoptera of more than 475,000 specimens acquired in 2015, when the MNCR received the collection from the National Institute of Biodiversity (INBio).
This collection contains a significant amount of type material corresponding to species described between the years 1901 and 2021. However, many of the paratypes from these publications were deposited in the Museum, but they were not systematized when incorporating the database system, therefore that there was not a complete inventory, until now.
The objective of this research was to generate the final inventory of the paratypes of the hymenoptera collection, through an exhaustive physical and literature review, in order to know the species and volume of specimens that are kept.
2. Methodology
2.1. National Museum Insect Collection
The MNCR paratype collection is found together with the general collection, along with the regular specimens. Although some of them held a label that identifies them as paratypes, they were not included in the digital database, so we proceeded to assign a code and enter their data into the system.
2.2. Paratype Inventory
By reviewing the literature, it was possible to build an inventory of specimens that was later confirmed by means of a physical search to count and confirm their presence in the collection. It was also necessary to have the documents inherited by the National Institute of Biodiversity (INBio) on the loans made, in order to locate the missing specimens in the collection.
2.3. Literature Review
We proceeded to carry out a review of the literature in order to avoid that some paratypes without a type label were not counted and identified in the database. For this we reviewed the following articles: Arcterberg et al. (2002) [1], Aranguren (1999) [2], Areekul and Quicke (2006) [3], Braet and Quicke (2004) [4], Briceño (2003) [5], Campra and Wahis (2018) [6], Carpenter and Kojima (2002) [7], Engel (2009) [8], Feitosa and Brandão (2008) [9], Fortier and Nishida (2004) [10], Gauld and Janzen (2004) [11], Gauld et al. (1997) [12], Gauld et al. (2000) [13], Gauld et al. (2002) [14], Gauld et al. (1998) [15], Gates and Hanson (2009) [16], Gibson (2003) [17], Gibson (2004) [18], Hansson (2005) [19], Hansson (2010) [20], Hansson (2011) [21], Hansson (2012) [22], Hansson and LaSalle (2003) [23], Heydon and Hanson (2005) [24], Leathers et al. (2003) [25], Lindsay and Sharkey (2006) [26], Longino (2006) [27], Macedo (2011) [28], Marsh (2002) [29], Marsh et al. (2013) [30], Michener et al. (2003) [31], Noyes (2004) [32], Noyes (2000) [33], Noyes and Hanson (1996) [34], Noyes and Ren (1995) [35], Noyes and Shauff (2003) [36], Roubik et al. (1997) [37], Sarmiento-Monroy (2006) [38], Schauff and Janzen (2001) [39], Sharkov and Wooley (1997) [40], Shaw et al. (1997) [41], Smith (2003) [42], Smith and Janzen (2003) [43], Smith (2008) [44], Smith (2012) [45], Starr (1991) [46], Twones (1992) [47], Valerio and Whitefield (2015) [48], Valerio and Whitefield (2003) [49], Valerio et al. (2009) [50], Vardy (2000) [51], Viereck (1920) [52], West-Eberhard et al. (2010) [53].
3. Results
The literature indicates that at least 4.218 paratypes are deposited in the MNCR and all of them were found (Table 1), and 65 have barcoding sequencing (Table 2). Unfortunately, some specimens returned damaged from a loan (Table 3), so
FAMILY |
Number of specimens in collection |
Subfamily |
tribe |
Species |
Specimens |
APIDADE |
|
|
Apinae |
|
|
Meliponini |
Meliwillea bivea Roubik, Lobo and Camargo 1997 |
2 specimens |
ARGIDAE |
|
|
Dielocerinae |
Themos mayi Smith and Janzen 2003 |
1 specimen |
Sterictiphorinae |
Didymia jonesi Smith 2003 |
1 specimen |
AULACIDAE |
|
|
Aulacinae |
|
|
|
Aulacus maculosus Smith 2008 |
1 specimen |
|
Pristaulacus nigricoxae Smith 2008 |
1 specimen |
BRACONIDAE |
|
|
Agatinae |
Alabagrus combos Leathers and Sharkey 2003 |
2 specimens |
|
Alabagrus derailersi Leathers and Sharkey 2003 |
8 specimens |
|
Alabagrus englishi Leathers and Sharkey 2003 |
43 specimens |
|
Alabagrus sarapiqui Leathers and Sharkey 2003 |
2 specimens |
|
Alabagrus watsoni Leathers and Sharkey 2003 |
5 specimens |
|
Amputoearinus niger Lindsay and Sharkey 2006 |
1 specimen |
|
Amputoearinus variegatus Lindsay and Sharkey 2006 |
3 specimens |
|
Sesioctonus acrolophus Briceño 2003 |
2 specimens |
|
Sesioctonus ariasi Briceño 2003 |
1 specimen |
|
Sesioctonus biospleres Briceño 2003 |
2 specimens |
|
Sesioctonus diazi Briceño 2003 |
1 specimen |
|
Sesioctonus dichromus Briceño 2003 |
11 specimens |
|
Sesioctonus miyayensis Briceño 2003 |
2 specimens |
|
Sesioctonus parathyridis Briceño 2003 |
5 specimens |
|
Zelomorpha bejaranoi Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha cherylae Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha euryterga Sarmiento-Monroy 2006 |
2 specimen |
|
Zelomorpha hallwachsae Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha janzeni Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha katatonus Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha miza Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha petila Sarmiento-Monroy 2006 |
1 specimen |
|
Zelomorpha sharkeyi Sarmiento-Monroy 2006 |
1 specimen |
Braconinae |
Compsobraconoides cinnamomi Fortier and Nishida 2004 |
3 specimens |
Doryctinae |
Barbalhoa contophleba Marsh 2002 |
2 specimens |
|
Barbalhoa licina Marsh 2002 |
13 specimens |
|
Curtiselloides pilosus Marsh 2002 |
20 specimens |
|
Notiospathius carolinae (Marsh 2002)a |
33 specimens |
|
Notiospathius pauli (Marsh 2002) |
4 specimens |
|
Iare belokobylskiji Marsh 2002 |
1 specimen |
|
Iare gauldi Marsh 2002 |
1 specimen |
|
Pannuceus fonsecai Marsh 2002 |
7 specimens |
|
Rimacollus rojasi Marsh 2002 |
1 specimen |
|
Ptesimogaster hespenheidi (Marsh 2002) |
5 specimens |
|
Tripteroides janzeni Marsh 2002 |
2 specimens |
Binareini |
Liobracon costaricensis Marsh 2002 |
3 specimens |
Doryctini |
Acanthorhogas parkeri Marsh 2002 |
1 specimen |
|
Gymnobracon megistus Marsh 2002 |
31 specimens |
|
Johnsonius costaricensis Marsh 2002 |
4 specimens |
|
Johnsonius perknus Marsh 2002 |
9 specimens |
|
Pedinotus (Pedinotus) niger Marsh 2002 |
2 specimens |
|
Pedinotus (Pedinotus) pardalotus Marsh 2002 |
3 specimens |
|
Pedinotus (Pedinotus) rojasi Marsh 2002 |
2 specimens |
|
Pedinotus (Pedinotus) variegatus Marsh 2002 |
1 specimen |
|
Rhaconotus (Pedinotus) costaricensis Marsh 2002 |
3 specimens |
|
Rhaconotus (Pedinotus) emarginatus Marsh 2002 |
14 specimens |
|
Rhaconotus (Pedinotus) rugosus Marsh 2002 |
1 specimen |
Ecphylini |
Bohartiellus costaricensis Marsh 2002 |
4 specimens |
|
Ecphylus (Ecphylus) gauldi Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) hansoni Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) hespenheidei Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) laselvus Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) leukocoxalis Marsh 2002 |
2 specimens |
|
Ecphylus (Ecphylus) lissoprotomus Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) propodealis Marsh 2002 |
1 specimen |
|
Ecphylus (Ecphylus) variabilis Marsh 2002 |
2 specimens |
Hecabolini |
Doryctinus costaricensis (Marsh 2002) |
6 specimens |
|
Doryctinus erugatus (Marsh 2002) |
1 specimen |
|
Doryctinus rubronotum (Marsh 2002) |
3 specimens |
|
Allorhogas ardisia Marsh 2002 |
1 specimen |
|
Allorhogas costaricensis Marsh 2002 |
2 specimens |
|
Allorhogas gauldi Marsh 2002 |
1 specimen |
|
Allorhogas hansoni Marsh 2002 |
1 specimen |
|
Allorhogas ingavera Marsh 2002 |
4 specimens |
|
Allorhogas sulcatus Marsh 2002 |
1 specimen |
|
Pioscelus costaricensis (Marsh 2002) |
3 specimens |
|
Curtisella gauldi Marsh 2002 |
2 specimens |
|
Hecabolus costaricensis Ugalde and Gauld 2002 |
1 specimen |
|
Heredius laselvus Marsh 2002 |
1 specimen |
|
Leluthia (Leluthia) careovena Marsh 2002 |
2 specimens |
|
Leluthia (Leluthia) costaricensis Marsh 2002 |
1 specimen |
|
Leluthia (Leluthia) flavocoxalis Marsh 2002 |
7 specimens |
|
Semirhytus hansoni Marsh 2002 |
16 specimens |
Heterospilini |
Pioscelus mesomphatus Marsh 2002 |
3 specimens |
|
Heterospilus (Heterospilus) alajuelus Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) aubreyae Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) barbaloae Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) carolinae Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) chaoi Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) corcovado Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) costarricensis Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) ektorincon Marsh 2013 |
3 specimens |
|
Heterospilus (Heterospilus) gouleti Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) granulatus Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) hansoni Marsh 2013 |
5 specimens |
|
Heterospilus (Heterospilus) jennieae Marsh 2013 |
26 specimens |
|
Heterospilus (Heterospilus) kellieae Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) leenderti Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) leviscutum Marsh 2013 |
3 specimens |
|
Heterospilus (Heterospilus) masneri Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) mercury Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) miskito Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) nahua Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) nephus Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) nueve Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) nunesi Marsh 2013 |
7 specimens |
|
Heterospilus (Heterospilus) orbitus Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) petralbus Marsh 2013 |
5 specimens |
|
Heterospilus (Heterospilus) rhabdotus Marsh 2013 |
4 specimens |
|
Heterospilus (Heterospilus) rincovensis Marsh 2013 |
3 specimens |
|
Heterospilus (Heterospilus) rojasi Marsh 2013 |
1 specimen |
|
Heterospilus (Heterospilus) sergeyi Marsh 2013 |
4 specimens |
|
Heterospilus (Heterospilus) shawi Marsh 2013 |
3 specimens |
|
Heterospilus (Heterospilus) spiloheterus Marsh 2013 |
2 specimens |
|
Heterospilus (Heterospilus) whartoni Marsh 2013 |
1 specimen |
Spathiini |
Notiospathius bicolor Marsh 2002 |
1 specimen |
|
Notiospathius rugonotum Marsh 2002 |
1 specimen |
|
Psenobolus triangularisvan Achterberg and Marsh 2002 |
3 specimens |
|
Spathius (Spathius) albocoxus Marsh 2002 |
18 specimens |
|
Trigonophasmus costaricensis Marsh 2002 |
4 specimens |
Mendesellinae |
Epsilogaster antoniae Valerio and Whitefield 2004 |
2 specimens |
Microgastrinae |
|
|
Microgastrini |
Hypomicrogaster koino Valerio 2015 |
3 specimens |
|
Hypomicrogaster largus Valerio 2015 |
1 specimen |
|
Hypomicrogaster luisi Valerio 2015 |
3 specimens |
|
Hypomicrogaster plagios Valerio 2015 |
10 specimens |
|
Hypomicrogaster rugosus Valerio 2015 |
3 specimens |
|
Hypomicrogaster specialis Valerio 2015 |
14 specimens |
|
Hypomicrogaster zan Valerio 2015 |
17 specimens |
|
Parapanteles continua Valerio and Whitefield 2009 |
11 specimens |
|
Parapanteles em Valerio and Whitefield 2009 |
7 specimens |
|
Parapanteles mariae Valerio and Whitefield 2009 |
15 specimens |
|
Parapanteles polus Valerio and Whitefield 2009 |
5 specimens |
|
Parapanteles rarus Valerio and Whitefield 2009 |
5 specimens |
|
Parapanteles scotti Valerio and Whitefield 2009 |
8 specimens |
|
Parapanteles sicpolus Valerio and Whitefield 2009 |
4 specimens |
|
Parapanteles tessares Valerio and Whitefield 2009 |
15 specimens |
|
Parapanteles tlinea Valerio and Whitefield 2009 |
5 specimens |
Orgilinae |
|
|
Mimagathidini |
Stantonia osa Braet and Quicke 2004 |
5 specimens |
|
Stantonia peninsularis Braet and Quicke 2004 |
1 specimen |
|
Stantonia sirena Braet and Quicke 2004 |
14 specimens |
Rogadinae |
|
|
Rogadini |
Aleiodes (Aleiodes) notozophus Marsh and Shaw 1997 |
2 specimens |
Yeliconini |
Yelicones limonensis Areekul and Quicke 2006 |
1 specimen |
|
Yelicones ormei Areekul and Quicke 2006 |
3 specimens |
|
Yelicones quarterus Areekul and Quicke 2006 |
1 specimen |
COLLETIDAE |
|
|
Diphaglossinae |
|
|
Caupolicanini |
|
|
|
Caupolicana inbio Michener, Engel and Ayala 2003 |
54 specimens |
|
Caupolicana rozenorum Michener, Engel and Ayala 2003 |
1 specimen |
ENCYRTIDAE |
|
|
Encyrtinae |
Bennettisca ganyame Noyes 2004 |
1 specimen |
|
Bennettisca nambue Noyes 2004 |
1 specimen |
|
Bennettisca xenophon Noyes 2004 |
1 specimen |
|
Blanchardiscus pollux Noyes 2004 |
3 specimens |
|
Caldencyrtus mitchelli Noyes and Hanson 1996 |
1 specimen |
|
Cercobelus sithon Noyes and Hanson 1996 |
1 specimen |
|
Cercobelus godoyae Noyes and Hanson 1996 |
1 specimen |
|
Cercobelus isara Noyes and Hanson 1996 |
2 specimens |
|
Metaphycus agylla Noyes 2004 |
1 specimen |
|
Metaphycus alsis Noyes 2004 |
4 specimens |
|
Metaphycus bostra Noyes 2004 |
1 specimen |
|
Metaphycus bouceki Noyes 2004 |
19 specimens |
|
Metaphycus carveri Noyes 2004 |
2 specimens |
|
Metaphycus ceros Noyes 2004 |
1 specimen |
|
Metaphycus chicle Noyes 2004 |
1 specimen |
|
Metaphycus cinna Noyes 2004 |
1 specimen |
|
Metaphycus codrus Noyes 2004 |
2 specimens |
|
Metaphycus coeus Noyes 2004 |
1 specimen |
|
Metaphycus coronea Noyes 2004 |
1 specimen |
|
Metaphycus cossyra Noyes 2004 |
4 specimens |
|
Metaphycus cures Noyes 2004 |
2 specimens |
|
Metaphycus cyrnos Noyes 2004 |
1 specimen |
|
Metaphycus cythera Noyes 2004 |
1 specimen |
|
Metaphycus dirce Noyes 2004 |
10 specimens |
|
Metaphycus electra Noyes 2004 |
1 specimen |
|
Metaphycus elisa Noyes 2004 |
2 specimens |
|
Metaphycus ephyrus Noyes 2004 |
2 specimens |
|
Metaphycus eruptor Noyes 2004 |
1 specimen |
|
Metaphycus erynis Noyes 2004 |
3 specimens |
|
Metaphycus eryx Noyes 2004 |
1 specimen |
|
Metaphycus expero Noyes 2004 |
1 specimen |
|
Metaphycus fasula Noyes 2004 |
2 specimen |
|
Metaphycus fautus Noyes 2004 |
3 specimens |
|
Metaphycus flavus Noyes 2004 |
12 specimens |
|
Metaphycus galata Noyes 2004 |
3 specimens |
|
Metaphycus gallicola Noyes and Hanson 1996 |
2 specimens |
|
Metaphycus genava Noyes 2004 |
2 specimens |
|
Metaphycus hamadra Noyes 2004 |
1 specimen |
|
Metaphycus harveyorum Noyes 2004 |
4 specimens |
|
Metaphycus hippo Noyes 2004 |
2 specimens |
|
Metaphycus inachus Noyes 2004 |
1 specimen |
|
Metaphycus lexova Noyes 2004 |
2 specimens |
|
Metaphycus lebadus Noyes 2004 |
1 specimen |
|
Metaphycus lucar Noyes 2004 |
1 specimen |
|
Metaphycus manda Noyes 2004 |
1 specimen |
|
Metaphycus masa Noyes 2004 |
1 specimen |
|
Metaphycus machaon Noyes 2004 |
1 specimen |
|
Metaphycus medio Noyes 2004 |
1 specimen |
|
Metaphycus memphis Noyes 2004 |
2 specimens |
|
Metaphycus merendo Noyes 2004 |
10 specimens |
|
Metaphycus metor Noyes 2004 |
15 specimens |
|
Metaphycus mellis Noyes 2004 |
1 specimen |
|
Metaphycus murena Noyes 2004 |
1 specimen |
|
Metaphycus mus Noyes 2004 |
1 specimen |
|
Metaphycus mycene Noyes 2004 |
2 specimens |
|
Metaphycus naxos Noyes 2004 |
3 specimens |
|
Metaphycus nola Noyes 2004 |
1 specimen |
|
Metaphycus norahae Noyes 2004 |
5 specimens |
|
Metaphycus obstinus Noyes 2004 |
1 specimen |
|
Metaphycus omega Noyes 2004 |
4 specimens |
|
Metaphycus othyras Noyes 2004 |
1 specimen |
|
Metaphycus paramo Noyes 2004 |
1 specimen |
|
Metaphycus porsena Noyes 2004 |
1 specimen |
|
Metaphycus portoricensis Noyes 2004 |
1 specimen |
|
Metaphycus pydnos Noyes 2004 |
3 specimens |
|
Metaphycus sestus Noyes 2004 |
1 specimen |
|
Metaphycus xsara Noyes 2004 |
1 specimen |
|
Paramucrona srunua Noyes 2004 |
1 specimen |
|
Pseudleptomastix mexicana Noyes and Schauff, 2003 |
2 specimens |
|
Psyllaephagus creusa Noyes and Hanson 1996 |
4 specimens |
|
Psyllaephagus epulo Noyes and Hanson 1996 |
1 specimen |
|
Psyllaephagus gyces Noyes and Hanson 1996 |
1 specimen |
|
Psyllaephagus othrys Noyes and Hanson 1996 |
2 specimens |
|
Psyllaephagus ufens Noyes and Hanson 1996 |
1 specimen |
Tetracneminae |
Acerophagoides almon Noyes 2000 |
5 specimens |
|
Acerophagoides cypris Noyes 2000 |
1 specimen |
|
Aenasius dives Noyes and Ren 1995 |
1 specimen |
|
Aenasius quezadai Noyes and Ren 1995 |
1 specimen |
|
Anagyrus cepio Noyes 2000 |
1 specimen |
|
Anagyrus matho Noyes 2000 |
1 specimen |
|
Anagyrus mirtesae Noyes 2000 |
2 specimens |
|
Anagyrus lizanorum Noyes 2000 |
15 specimens |
|
Anagyrus odacon Noyes 2000 |
1 specimen |
|
Anagyrus remotor Noyes, 2000 |
1 specimen |
|
Anagyrus salazari Noyes 2000 |
1 specimen |
|
Anagyrus sucro Noyes 2000 |
2 specimens |
|
Anagyrus villalobosi Noyes 2000 |
6 specimens |
|
Blepyrus fabius Noyes 2000 |
4 specimens |
|
Blepyrus hansoni Noyes 2000 |
25 specimens |
|
Blepyrus sappho Noyes 2000 |
7 specimens |
|
Blepyrus zenonis Noyes 2000 |
6 specimens |
|
Chrysoplatycerus ixion Noyes 2000 |
1 specimen |
|
Clausenia senex Noyes 2000 |
2 specimens |
|
Gyranusoidea amasis Noyes 2000 |
9 specimens |
|
Gyranusoidea brixia Noyes 2000 |
1 specimen |
|
Gyranusoidea sulmo Noyes 2000 |
3 specimens |
|
Gyranusoidea janzeni Noyes 2000 |
1 specimen |
|
Gyranusoidea lamus Noyes 2000 |
4 specimens |
|
Gyranusoidea rhodope Noyes 2000 |
10 specimens |
|
Hambletonia pilosifrons Sharkov and Woolley 1997 |
2 specimens |
|
Hipponactis itea Noyes 2000 |
9 specimens |
|
Peneax deidamia Noyes 2000 |
2 specimens |
|
Zarhopalus zancles Noyes 2000 |
6 specimens |
EULOPHIDAE |
|
|
Entedoninae |
|
|
|
Achrysocharoides infuscus Hansson, 2012 |
1 specimen |
|
Aleuroctonus marki Hansson and LaSalle 2003 |
1 specimen |
|
Apleurotropis albicaulis Hansson 2010 |
12 specimens |
|
Apleurotropis albiscapus Hansson 2010 |
3 specimens |
|
Apleurotropis anemia Hansson 2010 |
24 specimens |
|
Apleurotropis assis Hansson 2010 |
3 specimens |
|
Cabeza baeostigma Hansson and LaSalle 2003 |
4 specimens |
|
Cabeza petiolata Hansson and LaSalle 2003 |
2 specimens |
|
Cornugon bicornis Hansson 2011 |
1 specimen |
|
Cornugon diabolos Hansson 2011 |
1 specimen |
|
Cornugon diceros Hansson 2011 |
1 specimen |
|
Cornugon gibberum Hansson 2011 |
1 specimen |
|
Cornugon leios Hansson 2011 |
2 specimens |
|
Cornugon reticulatum Hansson 2011 |
5 specimens |
|
Dentalion alveum Hansson 2011 |
1 specimen |
|
Dentalion apon Hansson 2011 |
1 specimen |
|
Klyngon jimenezi Hansson 2005 |
4 specimens |
|
Klyngon serjaniae Hansson and LaSalle, 2010 |
1 specimen |
Eulophinae |
|
|
|
Euplectrus mariae Schauff 2001 |
8 specimens |
|
Euplectrus xiomarae Schauff 2001 |
5 specimens |
EURYTOMIDAE |
|
|
Eurytominae |
Bephrata camacho Gates and Hanson 2009 |
7 specimens |
|
Bephrata flava Gates and Hanson 2009 |
2 specimens |
|
Bephrata lorraineae Gates and Hanson 2009 |
7 specimens |
|
Bephrata nublada Gates and Hanson 2009 |
1 specimen |
|
Bephrata stichogaster Gates and Hanson 2009 |
6 specimens |
|
Bephrata ticos Gates and Hanson 2009 |
17 specimens |
|
Isosomodes azofiefai Gates and Hanson 2009 |
19 specimens |
FORMICIDAE |
|
|
Myrmicinae |
|
|
Dacetini |
Nesomyrmex vargasi Longino 2006 |
1 specimen |
|
Pyramica oconitrilloae Longino 2006 |
1 specimen |
|
Pyramica paniaguae Longino 2006 |
5 specimens |
Attini |
Lachnomyrmex laticeps Feitosa and Brandao 2008 |
1 specimen |
|
Lachnomyrmex longinoi Feitosa and Brandao 2008 |
3 specimens |
|
Lachnomyrmex regularis Feitosa and Brandao 2008 |
3 specimens |
|
Strumigenys paniaguae (Longino 2006) |
1 specimen |
GASTERUPTIIDAE |
|
|
Gasteruptiinae |
Gasteruption hansoni Macedo 2011 |
4 specimens |
HALICTIDAE |
|
|
Halictinae |
|
|
Augochlorini |
Megalopta atra Engel 2006 |
21 specimens |
ICHNEUMONIDAE |
|
|
Anomaloninae |
|
|
Anomalonini |
Anomalon acevedoi Gauld 1997 |
27 specimens |
|
Anomalon anae Gauld and Bradshaw 1997 |
8 specimens |
|
Anomalon avanzoi Gauld 1997 |
4 specimens |
|
Anomalon canoae Gauld 1997 |
23 specimens |
|
Anomalon fermini Gauld 1997 |
10 specimens |
|
Anomalon guisellea Gauld 1997 |
8 specimens |
Gravenhorstiini |
Castrosion renei Gauld 1997 |
2 specimens |
|
Corsoncus minori Gauld 1997 |
1 specimen |
|
Podogaster acostai Gauld and Bradshaw 1997 |
2 specimens |
|
Podogaster benignoi Gauld and Bradshaw 1997 |
1 specimen |
|
Podogaster eldae Gauld and Bradshaw 1997 |
12 specimens |
|
Podogaster orozcoi Gauld 1997 |
1 specimen |
|
Podogaster tranae Gauld and Bradshaw 1997 |
10 specimens |
Banchinae |
|
|
Atrophini |
Cordeleboea nasuta Ugalde and Gauld 2002 |
2 specimens |
|
Cordeleboea nigrata Ugalde and Gauld 2002 |
1 specimen |
|
Cordeleboea ovardesi Ugalde and Gauld 2002 |
1 specimen |
|
Hadrostethus eberhardi Ugalde and Gauld 2002 |
12 specimens |
|
Hadrostethus hansoni Ugalde and Gauld 2002 |
2 specimens |
|
Hapsinotus albomaculatus Ugalde and Gauld 2002 |
9 specimens |
|
Hapsinotus bravicaudus Ugalde and Gauld 2002 |
2 specimens |
|
Hapsinotus carinatus Ugalde and Gauld 2002 |
20 specimens |
|
Hapsinotus convexus Ugalde and Gauld 2002 |
9 specimens |
|
Hapsinotus glissatus Ugalde and Gauld 2002 |
9 specimens |
|
Hapsinotus granulatus Ugalde and Gauld 2002 |
1 specimen |
|
Hapsinotus morenus Ugalde and Gauld 2002 |
4 specimens |
|
Hapsinotus nigrogena Ugalde and Gauld 2002 |
14 specimens |
|
Hapsinotus notaulator Ugalde and Gauld 2002 |
7 specimens |
|
Hapsinotus parvatus Ugalde and Gauld 2002 |
9 specimens |
|
Hapsinotus planifacies Ugalde and Gauld 2002 |
1 specimen |
|
Hapsinotus transversus Ugalde and Gauld 2002 |
1 specimen |
|
Lissonota begaza Ugalde and Gauld 2002 |
4 specimens |
|
Lissonota conulla Ugalde and Gauld 2002 |
29 specimens |
|
Lissonota crudeta Ugalde and Gauld 2002 |
14 specimens |
|
Lissonota gurna Ugalde and Gauld 2002 |
5 specimens |
|
Lissonota horpa Ugalde and Gauld 2002 |
1 specimen |
|
Lissonota justina Ugalde and Gauld 2002 |
1 specimen |
|
Lissonota pseudeleboea Ugalde and Gauld 2002 |
15 specimens |
|
Lissonota tostada Ugalde and Gauld 2002 |
2 specimens |
|
Lissonota vebena Ugalde and Gauld 2002 |
2 specimens |
|
Loxodocus luteator Ugalde and Gauld 2002 |
3 specimens |
|
Loxodocus nigratus Ugalde and Gauld 2002 |
7 specimens |
|
Loxodocus vulgator Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha alhambra Ugalde and Gauld 2002 |
3 specimens |
|
Meniscomorpha astrema Ugalde and Gauld 2002 |
28 specimens |
|
Meniscomorpha bullata Ugalde and Gauld 2002 |
3 specimens |
|
Meniscomorpha castroi Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha devisa Ugalde and Gauld 2002 |
10 specimens |
|
Meniscomorpha enura Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha gomezi Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha helspina Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha hembrata Ugalde and Gauld 2002 |
10 specimens |
|
Meniscomorpha implaza Ugalde and Gauld 2002 |
1 specimen |
|
Meniscomorpha leva Ugalde and Gauld 2002 |
1 specimen |
|
Meniscomorpha lizanoi Ugalde and Gauld 2002 |
1 specimen |
|
Meniscomorpha mitica Ugalde and Gauld 2002 |
5 specimens |
|
Meniscomorpha nerda Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha ocala Ugalde and Gauld 2002 |
6 specimens |
|
Meniscomorpha pensita Ugalde and Gauld 2002 |
3 specimens |
|
Meniscomorpha quava Ugalde and Gauld 2002 |
6 specimens |
|
Meniscomorpha romata Ugalde and Gauld 2002 |
9 specimens |
|
Meniscomorpha salasi Ugalde and Gauld 2002 |
1 specimen |
|
Meniscomorpha sotoi Ugalde and Gauld 2002 |
19 specimens |
|
Meniscomorpha tamaska Ugalde and Gauld 2002 |
2 specimens |
|
Meniscomorpha xeresha Ugalde and Gauld 2002 |
6 specimens |
|
Meniscomorpha zacasta Ugalde and Gauld 2002 |
4 specimens |
|
Mnioes arves Ugalde and Gauld 2002 |
2 specimens |
|
Mnioes cerves Ugalde and Gauld 2002 |
1 specimen |
|
Mnioes dormes Ugalde and Gauld 2002 |
2 specimens |
|
Mnioes flomes Ugalde and Gauld 2002 |
1 specimen |
|
Mnioes garnes Ugalde and Gauld 2002 |
14 specimens |
|
Mnioes kebes Ugalde and Gauld 2002 |
3 specimens |
|
Mnioes mores Ugalde and Gauld 2002 |
6 specimens |
|
Mnioes nalbes Ugalde and Gauld 2002 |
14 specimens |
|
Mnioes teves Ugalde and Gauld 2002 |
13 specimens |
|
Occia jereza Ugalde and Gauld 2002 |
9 specimens |
|
Podeleboea altissima Ugalde and Gauld 2002 |
7 specimens |
|
Podeleboea bifenestralis Ugalde and Gauld 2002 |
2 specimens |
|
Podeleboea bricenoi Ugalde and Gauld 2002 |
2 specimens |
|
Podeleboea cacaoi Ugalde and Gauld 2002 |
7 specimens |
|
Podeleboea morena Ugalde and Gauld 2002 |
1 specimen |
|
Podeleboea perplexa Ugalde and Gauld 2002 |
6 specimens |
|
Podeleboea pitillae Ugalde and Gauld 2002 |
1 specimen |
|
Podeleboea punctiblanca Ugalde and Gauld 2002 |
1 specimen |
|
Podeleboea vitoi Ugalde and Gauld 2002 |
4 specimens |
|
Procestus pammae Ugalde and Gauld 2002 |
3 specimens |
|
Ptychopsis glabrata Ugalde and Gauld 2002 |
1 specimen |
|
Ptychopsis sanchoi Ugalde and Gauld 2002 |
2 specimens |
|
Quillonota petronae Ugalde and Gauld 2002 |
1 specimen |
|
Quillonota zurqui Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus abonis Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus besamis Ugalde and Gauld 2002 |
11 specimens |
|
Syzeuctus cenitus Ugalde and Gauld 2002 |
2 specimens |
|
Syzeuctus clovis Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus dorcus Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus forpis Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus hezonicus Ugalde and Gauld 2002 |
47 specimens |
|
Syzeuctus judtitus Ugalde and Gauld 2002 |
4 specimens |
|
Syzeuctus kespius Ugalde and Gauld 2002 |
1 specimen |
|
Syzeuctus lurinus Ugalde and Gauld 2002 |
2 specimens |
|
Syzeuctus possitus Ugalde and Gauld 2002 |
9 specimens |
|
Syzeuctus vedoris Ugalde and Gauld 2002 |
4 specimens |
|
Wahlamia anasiphaga Ugalde and Gauld 2002 |
1 specimen |
|
Wahlamia brevigena Ugalde and Gauld 2002 |
1 specimen |
|
Wahlamia illluminata Ugalde and Gauld 2002 |
2 specimens |
|
Wahlamia incognita Ugalde and Gauld 2002 |
5 specimens |
|
Wahlamia incompleta Ugalde and Gauld 2002 |
1 specimen |
|
Wahlamia phloridiphobia Ugalde and Gauld 2002 |
3 specimens |
|
Wahlamia taliskiphila Ugalde and Gauld 2002 |
11 specimens |
Banchini |
Exetastes amancoi Ugalde and Gauld 2002 |
1 specimen |
|
Exetastes eithus Ugalde and Gauld 2002 |
3 specimens |
|
Exetastes lorinus Ugalde and Gauld 2002 |
1 specimen |
|
Exetastes navetus Ugalde and Gauld 2002 |
1 specimen |
|
Exetastes niblus Ugalde and Gauld 2002 |
1 specimen |
|
Exetastes ometus Ugalde and Gauld 2002 |
10 specimens |
|
Exetastes pivai Ugalde and Gauld 2002 |
8 specimens |
|
Exetastes quarus Ugalde and Gauld 2002 |
20 specimens |
Glyptini |
Glypta eberhardi Ugalde and Gauld 2002 |
1 specimen |
|
Glypta metadecoris Ugalde and Gauld 2002 |
1 specimen |
|
Glypta vittata Ugalde and Gauld 2002 |
1 specimen |
|
Glypta wahli Ugalde and Gauld 2002 |
1 specimen |
|
Glypta zurquiensis Ugalde and Gauld 2002 |
1 specimen |
|
Sphelodon ugaldei Ugalde and Gauld 2002 |
1 specimen |
|
Sphelodon wardae Ugalde and Gauld 2002 |
3 specimens |
|
Zaglytomorpha auxiliadorae Ugalde and Gauld 2002 |
2 specimens |
|
Zaglytomorpha bella Ugalde and Gauld 2002 |
1 specimen |
|
Zaglytomorpha cornuta Ugalde and Gauld 2002 |
1 specimen |
|
Zaglytomorpha daschi Ugalde and Gauld 2002 |
1 specimen |
|
Zaglytomorpha flaquis Ugalde and Gauld 2002 |
4 specimens |
|
Zaglytomorpha mitchellae Ugalde and Gauld 2002 |
5 specimens |
Cremastinae |
Eiphosoma gollum Gauld 2000 |
3 specimens |
|
Eiphosoma porrax Gauld 2000 |
1 specimen |
|
Eiphosoma rudelium Gauld 1991 |
3 specimens |
|
Eiphosoma tantalium Gauld 1991 |
8 specimens |
|
Eiphosoma terrax Gauld 2000 |
2 specimens |
|
Eiphosoma urgulium Gauld 1991 |
4 specimens |
|
Pristomerus chicliphilus Gauld 2000 |
2 specimens |
|
Pristomerus grendor Gauld 2000 |
1 specimen |
|
Pristomerus haptinus Gauld 2000 |
1 specimen |
|
Pristomerus jugulator Gauld 2000 |
5 specimens |
|
Pristomerus kirinus Gauld 2000 |
2 specimens |
|
Ptilobaptus solaniphilus Gauld 2000 |
1 specimen |
|
Temelucha dannix Gauld 2000 |
37 specimens |
|
Trathala abraxa Gauld 2000 |
4 specimens |
|
Trathala celena Gauld 2000 |
8 specimens |
|
Trathala diva Gauld 2000 |
35 specimens |
|
Trathala gifa Gauld 2000 |
2 specimens |
|
Trathala kula Gauld 2000 |
3 specimens |
|
Xiphosomella belinda Gauld 2000 |
1 specimen |
|
Xiphosomella bonera Gauld 2000 |
51 specimens |
|
Xiphosomella cinctata Gauld 2000 |
28 specimens |
|
Xiphosomella ctenator Gauld 2000 |
10 specimens |
|
Xiphosomella daphne Gauld 2000 |
3 specimens |
|
Xiphosomella denticulella Gauld 2000 |
2 specimens |
|
Xiphosomella folicana Gauld 2000 |
1 specimen |
|
Xiphosomella forgana Gauld 2000 |
1 specimen |
|
Xiphosomella gethne Gauld 2000 |
1 specimen |
|
Xiphosomella hippolita Gauld 2000 |
6 specimens |
|
Xiphosomella jacusme Gauld 2000 |
12 specimens |
|
Xiphosomella kismeta Gauld 2000 |
33 specimens |
|
Xiphosomella merida Gauld 2000 |
1 specimen |
|
Xiphosomella ozne Gauld 2000 |
8 specimens |
|
Xiphosomella pesada Gauld 2000 |
1 specimen |
|
Xiphosomella pirena Gauld 2000 |
1 specimen |
|
Xiphosomella quinta Gauld 2000 |
1 specimen |
|
Xiphosomella roxana Gauld 2000 |
6 specimens |
|
Xiphosomella rufina Gauld 2000 |
1 specimen |
|
Xiphosomella slenda Gauld 2000 |
4 specimens |
|
Xiphosomella wirra Gauld 2000 |
3 specimens |
|
Xiphosomella zilla Gauld 2000 |
6 specimens |
Ctenopelmatinae |
|
|
Scolobatini |
Physotarsus bonillai Gauld 1997 |
2 specimens |
Cylloceriinae |
Cylloceria alvaradoi Gauld 1991 |
5 specimens |
|
Cylloceria barbouraki Gauld 1991 |
1 specimen |
|
Cylloceria trishae Gauld 1991 |
1 specimen |
|
Cylloceria ugaldevi Gauld 1991 |
1 specimen |
Labeninae |
|
|
Labenini |
Apechoneura compacta Gauld 2000 |
1 specimen |
|
Apechoneura mariae Gauld 2000 |
1 specimen |
|
Apechoneura pelucida Gauld 2000 |
1 specimen |
|
Apechoneura salablancae Gauld 2000 |
1 specimen |
|
Apechoneura valerieae Gauld 2000 |
4 specimens |
|
Labena eremica Gauld 2000 |
3 specimens |
|
Labena espinita Gauld 2000 |
1 specimen |
|
Labena humida Gauld 2000 |
1 specimen |
|
Labena ogra Gauld 2000 |
1 specimen |
|
Labena osai Gauld 2000 |
1 specimen |
Metopiinae |
Colpotrochia beluga Gauld and Sithole 2002 |
106 specimens |
|
Colpotrochia diabella Gauld and Sithole 2002 |
50 specimens |
|
Colpotrochia feroza Gauld and Sithole 2002 |
11 specimens |
|
Colpotrochia neblina Gauld and Sithole 2002 |
6 specimens |
|
Colpotrochia orinella Gauld and Sithole 2002 |
5 specimens |
|
Colpotrochia pesada Gauld and Sithole 2002 |
1 specimens |
|
Colpotrochia watanka Gauld and Sithole 2002 |
3 specimens |
|
Colpotrochia xenia Gauld and Sithole 2002 |
1 specimens |
|
Colpotrochia zembla Gauld and Sithole 2002 |
2 specimens |
|
Exochus boxatus Gauld and Ugalde 2002 |
2 specimens |
|
Exochus curvinus Gauld and Ugalde 2002 |
2 specimens |
|
Exochus delopius Gauld and Ugalde 2002 |
5 specimens |
|
Exochus evitus Gauld and Ugalde 2002 |
1 specimens |
|
Exochus farmellus Gauld and Ugalde 2002 |
6 specimens |
|
Exochus flubinus Gauld and Ugalde 2002 |
23 specimens |
|
Exochus gascus Gauld and Ugalde 2002 |
4 specimens |
|
Exochus hormus Gauld and Ugalde 2002 |
6 specimens |
|
Exochus izbus Gauld and Ugalde 2002 |
416 specimens |
|
Exochus jacintus Gauld and Ugalde 2002 |
14 specimens |
|
Exochus krellus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus nativus Gauld and Ugalde 2002 |
19 specimens |
|
Exochus nubosus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus ozanus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus pubitus Gauld and Ugalde 2002 |
3 specimens |
|
Exochus ravetus Gauld and Sithole 2002 |
13 specimens |
|
Exochus rubellus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus serozus Gauld and Ugalde 2002 |
11 specimens |
|
Exochus teborus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus urzus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus vanitus Gauld and Ugalde 2002 |
2 specimens |
|
Exochus vlops Gauld and Ugalde 2002 |
1 specimen |
|
Exochus voxanus Gauld and Ugalde 2002 |
3 specimens |
|
Exochus wabitus Gauld and Ugalde 2002 |
1 specimen |
|
Exochus womus Gauld and Ugalde 2002 |
20 specimens |
|
Exochus xarus Gauld and Ugalde 2002 |
23 specimens |
|
Exochus xetus Gauld and Ugalde 2002 |
6 specimens |
|
Exochus yorizus Gauld and Sithole 2002 |
6 specimens |
|
Forrestopius pamelae Gauld and Sithole 2002 |
2 specimens |
|
Hypsicera chena Gauld and Sithole 2002 |
11 specimens |
|
Hypsicera msana Gauld and Sithole 2002 |
1 specimen |
|
Hypsicera mukanwa Gauld and Sithole 2002 |
25 specimens |
|
Hypsicera nhema Gauld and Sithole 2002 |
1 specimen |
|
Hypsicera pfumbua Gauld and Sithole 2002 |
2 specimens |
|
Hypsicera ruokoa Gauld and Sithole 2002 |
9 specimens |
|
Leurus discus Gauld and Sithole 2002 |
6 specimens |
|
Leurus eraticus Gauld and Sithole 2002 |
5 specimens |
|
Leurus fascialis Gauld and Sithole 2002 |
5 specimens |
|
Leurus sylvaticus Gauld and Sithole 2002 |
1 specimen |
|
Metopius amenus Gauld and Sithole 2002 |
3 specimens |
|
Metopius buscus Gauld and Sithole 2002 |
6 specimens |
|
Metopius geophagus Gauld and Sithole 2002 |
8 specimens |
|
Metopius halorus Gauld and Sithole 2002 |
97 specimens |
|
Metopius pelus Gauld and Sithole 2002 |
6 specimens |
|
Seticornuta altamirae Gauld and Sithole 2002 |
1 specimen |
|
Stethoncus auberti Gauld and Sithole 2002 |
1 specimen |
|
Triclistus pailas Gauld and Sithole 2002 |
2 specimens |
|
Triclistus slimellus Gauld and Sithole 2002 |
7 specimens |
|
Triclistus tabetus Gauld and Sithole 2002 |
3 specimens |
|
Triclistus vaxinus Gauld and Sithole 2002 |
2 specimens |
|
Triclistus xodius Gauld and Sithole 2002 |
47 specimens |
|
Trieces azipas Gauld and Sithole 2002 |
3 specimens |
|
Trieces horisme Gauld and Sithole 2002 |
3 specimens |
|
Trieces riodinis Gauld and Sithole 2002 |
71 specimens |
|
Trieces tuvule Gauld and Sithole 2002 |
3 specimens |
|
Trieces wascia Gauld and Sithole 2002 |
51 specimens |
Ophioninae |
|
|
Thyreodonini |
Thyreodon darlingi Gauld 2004 |
1 specimen |
|
Thyreodon deansi Gauld 2004 |
1 specimen |
|
Thyreodon delvarei Gauld 2004 |
1 specimen |
|
Thyreodon papei Gauld 2004 |
1 specimen |
|
Thyreodon whitfieldi Gauld 2004 |
3 specimens |
|
Thyreodon zitaniae Gauld 2004 |
1 specimen |
Oxytorinae |
Oxytorus knappae Gauld and Mallet 2000 |
8 specimens |
Pimplinae |
|
|
Atrophini |
Anastelgis alturas Gates and Hanson 2009 |
1 specimen |
|
Anastelgis barvae Gauld, Ugalde and Hanson 1998 |
1 specimen |
|
Anastelgis garciai Gauld 1991 |
14 specimens |
|
Anastelgis jaramilloi Gauld 1991 |
3 specimens |
|
Anastelgis marini Gauld 1991 |
1 specimens |
Ephialtini |
Acrotaphus franklini Gauld 1991 |
3 specimens |
|
Calliephialtes sittenfeldae Gauld, Ugalde and Hanson 1998 |
1 specimen |
|
Clistopyga calixtoi Gauld 1991 |
11 specimens |
|
Clistopyga eldae Gauld 1991 |
9 specimens |
|
Clistopyga fernandezi Gauld 1991 |
8 specimens |
|
Clistopyga henryi Gauld 1991 |
10 specimens |
|
Dolichomitus cantillanoi Gauld 1991 |
3 specimens |
|
Dolichomitus flacissimus Gauld, Ugalde and Hanson 1998 |
1 specimen |
|
Dreisbachia dotae Hanson 1998 |
1 specimen |
|
Eruga gutfreundi Gauld 1991 |
2 specimens |
|
Eruga kauli Gauld 1991 |
1 specimens |
|
Eruga yehi Gauld 1991 |
2 specimens |
|
Hymenoepimecis robertsae Gauld 1991 |
1 specimen |
|
Iseropus barqueroi Gauld 1991 |
5 specimens |
|
Polysphincta dizardi Gauld 1991 |
1 specimen |
|
Polysphincta purcelli Gauld 1991 |
28 specimens |
|
Polysphincta shabui Gauld 1991 |
1 specimen |
|
Scambus basseyi Gauld 1991 |
3 specimens |
|
Tromatobia blancoi Gauld 1991 |
20 specimens |
|
Zatypota petronae Gauld 1991 |
7 specimens |
|
Zatypota solanoi Gauld 1991 |
1 specimen |
|
Zaglyptus chavesi Gauld 1991 |
28 specimens |
|
Zonopimpla barbosai Gauld 1991 |
3 specimens |
|
Zonopimpla duviliae Gauld 1991 |
1 specimen |
|
Zonopimpla hebeae Gauld 1991 |
1 specimen |
|
Zonopimpla lilae Gauld 1991 |
1 specimen |
|
Zonopimpla mirandai Gauld 1991 |
7 specimens |
|
Zonopimpla tenorioi Gauld 1991 |
1 specimen |
|
Zonopimpla zamorae Gauld 1991 |
1 specimen |
Pimplini |
Neotheronia abramsae Gauld 1991 |
38 specimens |
|
Neotheronia bostrandae Gauld 1991 |
5 specimens |
|
Neotheronia charli Gauld 1991 |
20 specimens |
|
Neotheronia donovani Gauld 1991 |
1 specimen |
|
Neotheronia holmae Gauld 1991 |
3 specimens |
|
Neotheronia jugaldei Gauld 1991 |
17 specimens |
|
Neotheronia kompassi Gauld 1991 |
1 specimen |
|
Neotheronia lizzae Gauld 1991 |
11 specimens |
|
Neotheronia lloydi Gauld 1991 |
3 specimens |
|
Neotheronia matamorosi Gauld 1991 |
39 specimens |
|
Neotheronia rosai Gauld 1991 |
42 specimens |
|
Nomosphecia godoyae Gauld, Ugalde and Hanson 1998 |
8 specimens |
|
Nomosphecia solisi Gauld 1991 |
24 specimens |
|
Pimpla rojasi Gauld 1991 |
16 specimens |
Poemeniinae |
|
|
Poemeniini |
Ganodes matai Gauld 1991 |
26 specimens |
Rhyssinae |
Epirhyssa corralesi Gauld 1991 |
3 specimens |
|
Epirhyssa curtisi Gauld 1991 |
1 specimen |
|
Epirhyssa zurquiae Gauld 1997 |
1 specimen |
Tryphoninae |
|
|
Oedemopsini |
Neliopisthus ocellatus Townes 1992 |
24 specimens |
|
Oedemopsis guemadoi Gauld 1997 |
2 specimens |
|
Zagryphus zulaya Gauld 1997 |
267 specimens |
Phytodietini |
Phytodietus penai Gauld 1997 |
8 specimens |
|
Phytodietus sylviae Gauld 1997 |
4 specimens |
|
Phytodietus valverdei Gauld 1997 |
2 specimens |
Sphinctini |
Sphinctus gastoni Gauld 1997 |
1 specimen |
Xoridinae |
Xoridess humos Gauld 1997 |
5 specimens |
|
Xorides secos Gauld 1997 |
6 specimens |
|
Xorides armidae Gauld 1997 |
2 specimens |
POMPILIDAE |
|
|
Pepsinae |
|
|
Pepsini |
Cryptocheilus neotropicalis Cambra and Wahis 2005 |
2 specimens |
|
Cryptocheilus santosi Cambra and Wahis 2005 |
10 specimens |
|
Pepsis inbio Vardy 2000 |
4 specimens |
PTEROMALIDAE |
|
|
Cleonyminae |
|
|
Lyciscini |
Shedoepistenia noyesi Gibson 2003 |
1 specimen |
Coelocybinae |
Cecidellis nigriseta Hanson 2005 |
12 specimens |
Spalangiinae |
Spalangia imitator Gibson 2009 |
2 specimens |
TENTHREDINIDAE |
|
|
Nematinae |
Pristiphora auricauda Smith 2003 |
1 specimen |
|
Pristiphora fuscalae Smith 2003 |
3 specimens |
TRIGONALYDAE |
Taeniogonalos woodorum Smith 2012 |
4 specimens |
VESPIDAE |
|
|
Polistinae |
|
|
Epiponini |
Chartergellus golfitensis West-Eberhard 2010 |
11 specimens |
|
Polybia bribri Carpenter 2002 |
30 specimens |
|
Polybia selvana Carpenter 2002 |
2 specimens |
a. According to zoological nomenclature, when a species changes gender, the author’s reference is placed in parentheses.
that only the pin with the labels remained, both specimens were glued directly to the pin. The paratypes represent 17 families, 41 subfamilies, 47 tribes and 143 genera.
4. Discussion
The paratype collection presents a significant loss of specimens, mostly due to old loans that were never returned, as well that 10 specimens of Heterospilus were lost during a loan process, only the empty pins with the labels arriving.
In the same way, some specimens were subjected to barcoding that were initially stored with the general collection, but because they now represent an additional resource, they are stored in a special collection of specimens with the same condition.
Museums usually represent a very important edge in scientific research, which not only has a merely taxonomic character of only naming new animal or plant species, but this knowledge impacts all other areas such as education [54], economy [55] and health [56]. This is why the maintenance of standard collections is vital, since technologies such as genome-wide sequencing require correctly identified specimens based on which to generate new information using their genetic material [57], and thus improve identification methods, and are the type specimens are those that will precisely be in the first row because they are the specimens from which a species is described.
Hence the need to improve the specimen loan processes, and the application of the laws on the deposit of duplicates and type specimens.
5. Conclusion
We conclude that all the specimens in the paratype collection are in perfect condition, with the exception of the Heterospilus specimens that were damaged during the return of material by mail. Likewise, the possibility of other paratypes re-entering loans that were never repaid remains on the table.
We are grateful to all colleagues who shared their publications or provided information to make this work possible. Likewise, thanks in advance to the research community, we are requesting their collaboration with the MNCR to locate the lost specimens, emphasizing that the collection that previously belonged to INBio is now in the MNCR, and the return of everything is required. The material is on loan, as many of the paratypes are found within that material and other colleagues need it for their research.