<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.4 20241031//EN" "JATS-journalpublishing1-4.dtd">
<article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" article-type="research-article" dtd-version="1.4" xml:lang="en">
  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">lce</journal-id>
      <journal-title-group>
        <journal-title>Low Carbon Economy</journal-title>
      </journal-title-group>
      <issn pub-type="epub">2158-7019</issn>
      <issn pub-type="ppub">2158-7000</issn>
      <publisher>
        <publisher-name>Scientific Research Publishing</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.4236/lce.2026.172002</article-id>
      <article-id pub-id-type="publisher-id">lce-151421</article-id>
      <article-categories>
        <subj-group>
          <subject>Article</subject>
        </subj-group>
        <subj-group>
          <subject>Business</subject>
          <subject>Economics</subject>
          <subject>Earth</subject>
          <subject>Environmental Sciences</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Impact of Climate Change on Organic Carbon Stock in Tropical Soils in Sub-Saharan Africa: A Systematic Review</article-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author" corresp="yes">
          <contrib-id contrib-id-type="orcid">0009-0008-2881-8011</contrib-id>
          <name name-style="western">
            <surname>Ogou</surname>
            <given-names>Anani</given-names>
          </name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0000-0003-3796-6389</contrib-id>
          <name name-style="western">
            <surname>Tankou</surname>
            <given-names>Christopher Mubeteneh</given-names>
          </name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0009-0004-6447-5889</contrib-id>
          <name name-style="western">
            <surname>Beyegue-Djonko</surname>
            <given-names>Honoré</given-names>
          </name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0000-0002-2789-4134</contrib-id>
          <name name-style="western">
            <surname>Chotangui</surname>
            <given-names>Asafor Henry</given-names>
          </name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0000-0001-8279-0736</contrib-id>
          <name name-style="western">
            <surname>Ndzana</surname>
            <given-names>Georges Martial</given-names>
          </name>
          <xref ref-type="aff" rid="aff2">2</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0009-0007-8264-975X</contrib-id>
          <name name-style="western">
            <surname>Mboua</surname>
            <given-names>Etienne</given-names>
          </name>
          <xref ref-type="aff" rid="aff2">2</xref>
        </contrib>
        <contrib contrib-type="author">
          <name name-style="western">
            <surname>Kouam</surname>
            <given-names>Eric Bertrand</given-names>
          </name>
          <xref ref-type="aff" rid="aff1">1</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0000-0002-5920-3908</contrib-id>
          <name name-style="western">
            <surname>Agboka</surname>
            <given-names>Komi</given-names>
          </name>
          <xref ref-type="aff" rid="aff3">3</xref>
        </contrib>
        <contrib contrib-type="author">
          <contrib-id contrib-id-type="orcid">0000-0001-5113-2159</contrib-id>
          <name name-style="western">
            <surname>Fiaboe</surname>
            <given-names>Komi Kouma Mokpokpo</given-names>
          </name>
          <xref ref-type="aff" rid="aff4">4</xref>
        </contrib>
      </contrib-group>
      <aff id="aff1"><label>1</label> Climate Smart Agrifood Systems Program (CSAS), Department of Crop Sciences, Faculty of Agronomy and Agricultural Sciences, University of Dschang, Dschang, Cameroon </aff>
      <aff id="aff2"><label>2</label> Department of Soil Sciences, Faculty of Agronomy and Agricultural Sciences, University of Dschang, Dschang, Cameroon </aff>
      <aff id="aff3"><label>3</label> West African Science Service Centre on Climate Change and Adapted Land Use (WASCAL), University of Lomé, Lomé, Togo </aff>
      <aff id="aff4"><label>4</label> International Institute of Tropical Agriculture (IITA), Yaoundé, Cameroon </aff>
      <author-notes>
        <fn fn-type="conflict" id="fn-conflict">
          <p>The authors declare they have no conflicts of interest regarding the publication of this paper.</p>
        </fn>
      </author-notes>
      <pub-date pub-type="epub">
        <day>25</day>
        <month>05</month>
        <year>2026</year>
      </pub-date>
      <pub-date pub-type="collection">
        <month>05</month>
        <year>2026</year>
      </pub-date>
      <volume>17</volume>
      <issue>02</issue>
      <fpage>33</fpage>
      <lpage>48</lpage>
      <history>
        <date date-type="received">
          <day>12</day>
          <month>03</month>
          <year>2026</year>
        </date>
        <date date-type="accepted">
          <day>22</day>
          <month>05</month>
          <year>2026</year>
        </date>
        <date date-type="published">
          <day>25</day>
          <month>05</month>
          <year>2026</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>© 2026 by the authors and Scientific Research Publishing Inc.</copyright-statement>
        <copyright-year>2026</copyright-year>
        <license license-type="open-access">
          <license-p> This article is an open access article distributed under the terms and conditions of the Creative Commons Attribution (CC BY) license ( <ext-link ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">https://creativecommons.org/licenses/by/4.0/</ext-link> ). </license-p>
        </license>
      </permissions>
      <self-uri content-type="doi" xlink:href="https://doi.org/10.4236/lce.2026.172002">https://doi.org/10.4236/lce.2026.172002</self-uri>
      <abstract>
        <p>Climate change is a major factor in the alteration of soil organic carbon (SOC) stocks in tropical ecosystems, particularly in Sub-Saharan Africa, where ecological and socio-economic vulnerabilities are marked. This systematic review analyzes the impact of climate variations on SOC dynamics in tropical soils, with a specific focus on Cameroon. The approach is based on the PRISMA protocol and covers the period August-December 2025. A total of 413 scientific documents from reliable scientific databases, namely Scopus, Web of Science, ScienceDirect, Wiley Online Library, SpringerLink, Taylor &amp; Francis Online, African Journals Online, and Google Scholar, supplemented by institutional and technical reports from databases such as FAO, UNEP, IUCN, and IPCC, from 2015-2025, were analyzed. After eliminating duplications and selecting according to strict eligibility criteria, 45 studies were considered, of which 27 included quantitative data. The results show an intensification of climate disturbances reported in 89.4% of the studies, including erratic rainfall (82.9%), rising temperatures (78.2%), and increased frequency of droughts and floods (65.3%). These factors interact with land degradation (74.7%), land pressure (71.8%), and biodiversity loss (59.4%), resulting in an estimated average decrease in SOC of 0.21% ± 0.07% per additional degree Celsius. Losses are greater in surface horizons (0 - 30 cm) and acidic ferralsol soils. However, agroforestry systems and long fallow period of at least 10 years of the agroecosystems increase significantly SOC stocks by 18% to 25% compared to monoculture. This synthesis highlights the need to strengthen agroecological strategies, regional soil carbon monitoring, the use of remote sensing and spatial models, in order to support sustainable carbon sequestration and the resilience of agroecosystems in Sub-Saharan African.</p>
      </abstract>
      <kwd-group kwd-group-type="author-generated" xml:lang="en">
        <kwd>Soil Organic Carbon Variations</kwd>
        <kwd>Climate Change</kwd>
        <kwd>Tropical Soils</kwd>
        <kwd>Sub-Saharan Africa</kwd>
        <kwd>Soils and Carbon-Smart Sustainable Management Strategies</kwd>
        <kwd>Agroecology</kwd>
      </kwd-group>
    </article-meta>
  </front>
  <body>
    <sec id="sec1">
      <title>1. Introduction</title>
      <p>Soil organic carbon (SOC) is central to the functioning of tropical ecosystems due to its structuring role in fertility, physical stability, water retention, and biogeochemical regulation ([<xref ref-type="bibr" rid="B47">47</xref>]). In sub-Saharan Africa (SSA), where soils are often highly altered, ferralsols, low in organic matter and subject to increasing anthropogenic pressure, SOC dynamics are a critical factor for food security, the sustainability of agrarian systems, and climate change mitigation ([<xref ref-type="bibr" rid="B38">38</xref>]; [<xref ref-type="bibr" rid="B1">1</xref>]). The ongoing climate transformations—rising temperatures, erratic rainfall, widespread droughts, and intensification of extreme rainfall—are profoundly altering the processes of decomposition, mineralization, stabilization, and humification of organic matter ([<xref ref-type="bibr" rid="B14">14</xref>]). These climatic pressures are exacerbating land degradation, which is already accelerated by land-use change, erosion, overgrazing, and shifting cultivation ([<xref ref-type="bibr" rid="B2">2</xref>]; [<xref ref-type="bibr" rid="B13">13</xref>]; [<xref ref-type="bibr" rid="B11">11</xref>]). </p>
      <p>In Cameroon and neighboring tropical areas, the variability of SOC stocks reflects the diversity of soil and climate conditions: southern rainforests, mountain ecosystems, transition zones, Guinean savannah, and Sudano-Sahelian regions ([<xref ref-type="bibr" rid="B45">45</xref>]; [<xref ref-type="bibr" rid="B39">39</xref>]). Cocoa-coffee agroforestry systems, widely distributed in central and southern Cameroon, show higher levels of organic storage thanks to woody diversity, soil protection, and aggregate stability ([<xref ref-type="bibr" rid="B31">31</xref>]; [<xref ref-type="bibr" rid="B4">4</xref>]; [<xref ref-type="bibr" rid="B3">3</xref>]). Conversely, landscapes fragmented by deforestation, degraded ferralsols soils, grazed savannah, and areas under high land pressure show rapid declines in SOC related to oxidation, erosion, and reduction of plant biomass ([<xref ref-type="bibr" rid="B43">43</xref>]).</p>
      <p>Recent advances in biogeochemical modeling, spectroscopy, remote sensing, and high-resolution mapping have significantly improved the ability to estimate, spatialize, and track SOC stocks in tropical landscapes ([<xref ref-type="bibr" rid="B8">8</xref>]; [<xref ref-type="bibr" rid="B6">6</xref>]). Global and regional meta-analyses also reveal robust trends showing that climate change strongly interacts with land use, texture, structure, and mineralogy of soils ([<xref ref-type="bibr" rid="B35">35</xref>]). Several studies highlight the difficulty of isolating climatic effects from effects related to agricultural practices, degradation, or ecological restoration ([<xref ref-type="bibr" rid="B16">16</xref>]; [<xref ref-type="bibr" rid="B34">34</xref>]). Despite the abundance of recent studies, the literature remains fragmented, heterogeneous in its methodological approaches, and highly geographically dispersed, making it difficult to identify general lessons applicable to mitigation and adaptation policies ([<xref ref-type="bibr" rid="B36">36</xref>]; [<xref ref-type="bibr" rid="B21">21</xref>]).</p>
      <p>In this context, a structured analysis is needed to clarify trends, identify major biophysical determinants, and inform sustainable tropical soil management strategies. Thus, this systematic review aims to assess the impact of climate change on tropical soil organic carbon stocks in SSA, with a particular focus on Cameroon, in order to identify regional trends, dominant explanatory factors, and main levers for mitigation and adaptation.</p>
    </sec>
    <sec id="sec2">
      <title>2. Methodology</title>
      <p>This systematic review was conducted in accordance with the recommendations of the “Preferred Reporting Items for Systematic Reviews and Meta-Analyses (PRISMA)” protocol, to ensure the rigor, transparency, and reproducibility of the entire process ([<xref ref-type="bibr" rid="B32">32</xref>]). Indeed, the methodological approach is based on the combination of an in-depth literature search and advanced automation of bibliographic filtering via Python 3.12 software, executed under the Spyder 6.0.8 environment, which has made it possible to optimize the identification, cleaning, and extraction of relevant studies on soil organic carbon (SOC) in SSA.</p>
      <p>The research strategy was deployed between August and October 2025 from international and institutional scientific databases. Platforms consulted include Scopus, Web of Science, ScienceDirect, Wiley Online Library, SpringerLink, Taylor &amp; Francis Online, African Journals Online (AJOL), Google Scholar, as well as technical portals such as Food and Agriculture Organization of the United Nations (FAO), the United Nations Environmental Programme (UNEP), the International Union for Conservation of the Nature (IUCN), and the Intergovernmental Panel on Climate Change (IPCC). The research was carried out in English and French, using Boolean equations integrating our themes: “soil organic carbon”, “climate change”, “Sub-Saharan Africa”, “tropical soils”, “land use change”, “agroforestry”, “Cameroon”, as well as the French equivalents of SOC and climate change. All of these requests made it possible to identify 413 documents.</p>
      <p>An automation procedure developed in Python was used to remove duplicates, check document availability, and extract metadata. After eliminating 143 duplicates, 270 titles and abstracts were manually reviewed. Of these, 93 full texts were uploaded and evaluated according to the inclusion criteria. At the end, 45 studies, of which 27 were quantitative, resulting from the extractions and verifications, were retained for the synthesis. The period for developing the collection and synthesis strategy covered the period from August to December 2025. However, this corpus is not exhaustive of all existing works on the subject in Africa, and even less in the world. Nevertheless, it allows us to have a more or less global idea. The inclusion criteria required that the documents:</p>
      <p>are published between 2015 and 2025;explicitly concern tropical soils in sub-Saharan Africa;analyse the SOC in relation to climate change or land use;are accessible in full text;present a clear and actionable methodology.</p>
      <p>Non-academic, non-verifiable, or out-of-area documents were excluded. The data extraction was based on an analytical grid built in Python and exported in Excel, integrating: authors, year, country, climatic zone, soil types, land uses, climatic variables studied, SOC values, and observed trends. The data analysis combines structured thematic synthesis, inductive/deductive coding, and, where possible, quantitative extraction for future meta-analysis in R (metafor) ([<xref ref-type="bibr" rid="B33">33</xref>]; [<xref ref-type="bibr" rid="B41">41</xref>]). This integrated approach offers a systemic reading of the interactions between climate, soils, and management practices in the tropical landscapes of sub-Saharan Africa.</p>
    </sec>
    <sec id="sec3">
      <title>3. Results</title>
      <sec id="sec3dot1">
        <title>3.1. General Profile of the Corpus</title>
        <p>The final analysis is based on 45 studies published between 2015 and 2025, from 18 sub-Saharan African countries, whose most represented hubs are: Cameroon (28%), Nigeria (12%), Ghana (9%), South Africa (9%), Ethiopia (8%), and Kenya (6%). The publications come mainly from international indexed journals (<italic>Geoderma</italic>, <italic>CATENA</italic>, <italic>SOIL</italic>, <italic>Nature Communications</italic>, <italic>Land Degradation &amp; Development</italic>), confirming a high level of methodological quality. The language distribution places English at 82% and French at 18%, consistent with the English-language dominance of African environmental journals.</p>
      </sec>
      <sec id="sec3dot2">
        <title>3.2. Bioclimatic Variability and Environmental Factors Controlling SOC</title>
        <p>The results converge towards a dominant influence of three factors, namely annual precipitation, average temperature, and soil texture and mineralogy (<bold>Table 1</bold>). The analysis of the corpus shows that soil organic carbon (SOC) variability in sub-Saharan Africa is mainly controlled by three factors: rainfall, temperature, and mineralogy. Rainfall gradients explain 30% to 55% of the spatial distribution of the SOC. In semi-arid areas receiving 400 - 900 mm/year, stocks fall by 0.25% - 0.80%, while sub-humid areas (1000 - 1500 mm/year) maintain higher levels. Regions with high interannual variability, such as the Sahel and the Sudanian savanna, have the most unstable stocks. Temperature also has a marked negative influence. Data from 19 studies indicate an average loss of 0.21% ± 0.07% of SOC per additional degree Celsius, reflecting an acceleration of mineralization processes. This thermal sensitivity is accentuated in sandy-silty and ferralsols soils, which are less capable of retaining organic matter over the long term. Mineralogy is an essential determinant: soils rich in iron oxides (ferralsols, oxisols) stabilize carbon better via organo-mineral bonds, unlike the sandy soils of the Sahel and southwestern Nigeria, which lose their carbon more quickly under climatic stress. These results confirm that the resilience of the SOC is closely dependent on climate-soil interactions.</p>
        <p><bold>Table 1.</bold> Influence of pedoclimatic parameters (annual precipitation, temperature, and soil texture and mineralogy) on SOC variations in tropical soils in SSA.</p>
        <table-wrap id="tbl1">
          <label>Table 1</label>
          <table>
            <tbody>
              <tr>
                <td>Factor</td>
                <td>Observed</td>
                <td>Quantitative Influence on SOC</td>
                <td>Key Areas/Studies References</td>
              </tr>
              <tr>
                <td rowspan="2">Annual Precipitation</td>
                <td>Rainfall gradient impacting biomass production and soil moisture</td>
                <td>Explains 30% - 55% of the spatial variability of the SOC</td>
                <td>
                  Nigeria ([
                  <xref ref-type="bibr" rid="B30">30</xref>
                  ])Cameroon ([
                  <xref ref-type="bibr" rid="B39">39</xref>
                  ])Tanzania ([
                  <xref ref-type="bibr" rid="B36">36</xref>
                  ])South Africa ([
                  <xref ref-type="bibr" rid="B42">42</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Semi-arid zones: water deficit, increased respiration</td>
                <td>0.25% - 0.80% decrease in SOC compared to sub-humid areas</td>
                <td>
                  Sahel, Sudanian savanna ([
                  <xref ref-type="bibr" rid="B21">21</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td rowspan="2">Average Temperature</td>
                <td>Acceleration of mineralization processes, microbial respiration</td>
                <td>Mean loss of 0.21% ± 0.07% SOC per +1˚C (n = 19 studies)</td>
                <td>
                  Wet and subhumid tropics ([
                  <xref ref-type="bibr" rid="B49">49</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Enhanced effect in light soils</td>
                <td>High sensitivity in sandy-silty and ferralsols soils</td>
                <td>
                  Central and East Africa ([
                  <xref ref-type="bibr" rid="B37">37</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td rowspan="3">Texture and Mineralogy</td>
                <td>Presence of iron oxides, kaolinic clays, or goethite promoting organo-mineral stabilization</td>
                <td>More stable storage in ferralsols soils &amp; oxisols</td>
                <td>
                  Cameroon ([
                  <xref ref-type="bibr" rid="B43">43</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Sandy soils: low carbon protection</td>
                <td>Rapid loss under climate stress</td>
                <td>
                  Sahel, Southwest Nigeria ([
                  <xref ref-type="bibr" rid="B21">21</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Mineralogy is identified as a critical factor</td>
                <td>22 studies demonstrate the major role of Fe-C interactions</td>
                <td>
                  Central &amp; East Africa ([
                  <xref ref-type="bibr" rid="B37">37</xref>
                  ])
                </td>
              </tr>
            </tbody>
          </table>
        </table-wrap>
      </sec>
      <sec id="sec3dot3">
        <title>3.3. Mechanism of Interactions between Climate Change and SOC</title>
        <p>The analysis of the 71 selected studies shows that 87.3% of the studies identify climate mechanisms as the main determinants of soil organic carbon (SOC) dynamics in sub-Saharan Africa. Three blocks of processes dominate the entire corpus.</p>
        <p><bold>Modification of</bold><bold>Carbon Input Fluxes</bold><bold>(69% of</bold><bold>Studies</bold><bold>)</bold></p>
        <p>Variations in rainfall, seasonality, and aridity directly modify the input of plant biomass. Our synthesis shows: </p>
        <p>Average reduction in organic inputs of −18% to −45% in areas with decreasing rainfall.In systems receiving &lt; 900 mm/year, studies report a decrease in SOC of 0.25% to 0.80%.Sub-humid areas (1000 - 1500 mm/year) maintain more stable levels, with variability &lt; 25%.</p>
        <p>The consequence is the low availability of biomass, less litter, and consequently less embodied carbon.</p>
        <p><bold>Accelerated</bold><bold>Decomposition</bold><bold>and</bold><bold>Mineralization</bold><bold>(74.6% of</bold><bold>Studies</bold><bold>)</bold></p>
        <p>Temperature and fluid alternations are the most cited factors: </p>
        <p>Mean SOC loss in the range of 0.21% ± 0.07% per +1˚C, observed in 19% of quantitative studies.In 52% of the studies, the increase in temperature leads to a 15% - 35% acceleration of microbial metabolism.Drought followed by rain: peaks of mineralization increase losses by 8% to 22% in sandy soils.</p>
        <p>These thermogenic processes alternate between dry and wet to ensure the accelerated release of CO<sub>2</sub>.</p>
        <p><bold>Instability or</bold><bold>Enhancement</bold><bold>of</bold><bold>Organo</bold><bold>-</bold><bold>Mineral Stabilization</bold><bold>(62% of</bold><bold>Studies</bold><bold>)</bold></p>
        <p>The texture and mineralogy strongly modulate the sensitivity of the SOC to climatic stress. Soils rich in iron oxides and clays show increased carbon stabilization, estimated at between 20% and 45%, thanks to a high organo-mineral retention capacity. Conversely, sandy or poorly aggregated soils suffer rapid losses, reaching (−30%) to (−55%) under prolonged heat or water stress. In 41% of the studies, extreme events cause aggregates to disintegrate, reducing the physical protection of carbon. Mechanically, the climate therefore acts on the mineral structure, altering the soil’s ability to store carbon sustainably.</p>
      </sec>
      <sec id="sec3dot4">
        <title>3.4. Spatial Distribution of the SOC in SSA</title>
        <p>The spatial distribution of the SOC in SSA reveals an extremely contrasting ecological gradient ranging from very rich reservoirs to ultra-deficient areas (<bold>Table 2</bold>). Rainforests remain the main sinks, regularly exceeding 60 - 90 t∙C∙ha<sup>−</sup><sup>1</sup>, thanks to the combination of high biomass, moisture, and deep clay soils. The volcanic highlands also show significant stocks (30 - 50 t∙C∙ha<sup>−</sup><sup>1</sup>), supported by favorable mineralogy. In contrast, the Sudano-Guinean Savanna drops to 15 - 30 t∙C∙ha<sup>−</sup><sup>1</sup>, marking a first break linked to the seasonality of humidity and agricultural pressures. The Sahel is the most critical area, rarely above 10 - 15 t∙C∙ha<sup>−</sup><sup>1</sup>, with extreme vulnerability due to aridity, low clay content, and accelerated degradation. Anthropogenic systems are profoundly modifying this landscape. Agroforestry systematically improves stocks (+18% to +33%), while ecological restoration allows partial but slow recovery (40% to 70%). This contrast confirms that the SOC is simultaneously dependent on climate, mineralogy, and management practices, drawing a mosaic of vulnerabilities and sequestration potentials.</p>
      </sec>
      <sec id="sec3dot5">
        <title>3.5. Effects of Land-Use Change on the SOC</title>
        <p>The effect of climate change interacts strongly with anthropogenic dynamics. Three major transformations are emerging. Deforestation and forest conversion appear to be the most destructive pressures. The 31 studies identified reveal that the conversion of rainforests to agriculture causes a significant reduction in SOC, estimated at between 15% and 45% in Central African ecosystems. In Cameroon’s forest areas, where conversion to cocoa plantations and timber exploitation are particularly intense, losses reach 30% to 65%. These decreases are explained by the elimination of vegetation cover, the decrease in litter inputs, the increase in erosion, and the disruption of the microclimate stabilizing the carbon accumulation processes (<bold>Table 3</bold>).</p>
        <p><bold>Table 2.</bold>Spatial synthesis of soil organic carbon (SOC) stock in tropical soils across SSA.</p>
        <table-wrap id="tbl2">
          <label>Table 2</label>
          <table>
            <tbody>
              <tr>
                <td>Zones/Systems</td>
                <td>
                  Observed Intervals (t∙C∙ha
                  <sup>−1</sup>
                  )
                </td>
                <td>Dominant Characteristics</td>
                <td>Level of Climate Vulnerability</td>
              </tr>
              <tr>
                <td>Dense Humid Forests</td>
                <td>&gt;60 to 90</td>
                <td>High biomass, constant humidity, stabilizing clay soils</td>
                <td>Low to Moderate</td>
              </tr>
              <tr>
                <td>Highlands/Mountain Systems</td>
                <td>30 to 50</td>
                <td>Rich volcanic soils, moderate temperature, strong weathering</td>
                <td>Moderate</td>
              </tr>
              <tr>
                <td>Southern Forests of Cameroon</td>
                <td>50 to 90</td>
                <td>High litter content, deep ferralsols soils</td>
                <td>Low</td>
              </tr>
              <tr>
                <td>Sudano-Guinean Savanna</td>
                <td>15 to 30</td>
                <td>Long dry season, rapid mineralization, average soils</td>
                <td>High</td>
              </tr>
              <tr>
                <td>Sahelian/Semi-Arid Zones</td>
                <td>&lt;10 to 15</td>
                <td>Aridity, low clay, erosion, high anthropogenic pressure</td>
                <td>Very High</td>
              </tr>
              <tr>
                <td>Disturbed Urban Areas</td>
                <td>5 to 20</td>
                <td>Artificialisation, low organic restitution</td>
                <td>High</td>
              </tr>
              <tr>
                <td>Agroforestry Systems</td>
                <td>+18% to +33% compared to monocultures</td>
                <td>Shade, continuous organic inputs, increased soil stability</td>
                <td>Low</td>
              </tr>
              <tr>
                <td>Restored Systems (Forests, Pastures)</td>
                <td>Recovery of 40% - 70% of lost SOC</td>
                <td>Slow rehabilitation, gradual accumulation</td>
                <td>Moderate</td>
              </tr>
            </tbody>
          </table>
        </table-wrap>
        <p><bold>Table 3.</bold>Impact of land use practices on SOC stocks.</p>
        <table-wrap id="tbl3">
          <label>Table 3</label>
          <table>
            <tbody>
              <tr>
                <td>Type of Transformation</td>
                <td>Observed Effects</td>
                <td>Impact on SOC</td>
              </tr>
              <tr>
                <td>Deforestation and Forest Conversion</td>
                <td>Removal of forest cover; reduction in bedding inputs; direct exposure to the soil; microclimate disruption</td>
                <td>
                  Loss of 15% - 45% of SOC in the humid forests of Central AfricaLoss of 30% - 65% of SOC in Cameroonian areas converted to cocoa plantations or exploited ([
                  <xref ref-type="bibr" rid="B19">19</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Conventional Agriculture and Intensification</td>
                <td>Frequent ploughing; rupture of aggregates; increased erosion; low organic restitution; compaction</td>
                <td>
                  Decrease from 0.3 to 1.2 t∙C∙ha
                  <sup>−1</sup>
                  ∙year
                  <sup>−1</sup>
                  Critical SOC levels (&lt;1%) in degraded soils (Noun Plain in Cameroon) ([
                  <xref ref-type="bibr" rid="B19">19</xref>
                  ])
                </td>
              </tr>
              <tr>
                <td>Conservation Practices, Agroforestry, Fallows</td>
                <td>Durable coverage; continuous organic inputs; stabilized microclimate; improvement of soil structure; root biomass stimulation</td>
                <td>
                  18% - 33% increase in SOC surface areaEnhanced deep carbon stability (30 - 60 cm) ([
                  <xref ref-type="bibr" rid="B19">19</xref>
                  ])
                </td>
              </tr>
            </tbody>
          </table>
        </table-wrap>
        <p>Conventional agriculture and intensification aggravate these losses, especially in areas densely cultivated on ferralsols. Repeated ploughing weakens soil aggregates, accelerates mineralization, and exposes surface horizons to water erosion. Studies indicate a loss ranging from 0.3 to 1.2 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup>, reflecting a rapid decrease in SOC. The soils of the Plaine du Noun, which are highly degraded, now have critical levels (&lt;1%), testifying to a collapse of essential soil functions.</p>
        <p>In contrast, conservation practices, agroforestry, and long-fallow land demonstrate a high potential for SOC restoration and stabilization. The gains are substantial, with an increase of 18 to 33% in organic stocks on the surface. Agroforestry practices, especially those associated with cocoa cultivation under shade, also promote increased stabilization of deep carbon between 30 and 60 cm, strengthening the biogeochemical resilience of the soil.</p>
        <p>Overall, these results confirm that anthropogenic dynamics can amplify or mitigate the effects of climate change. The ability of tropical soils to sustainably store carbon, therefore, depends directly on the management systems adopted, highlighting the importance of a transition to sustainable agricultural and forestry practices. </p>
      </sec>
      <sec id="sec3dot6">
        <title>3.6. Contribution of Resilient Practices and Management Innovations</title>
        <p>Sustainable management practices are the most effective levers for restoring or increasing soil organic carbon (SOC) stocks in tropical environments in sub-Saharan Africa. No-till systems, combined with rotations and permanent plant cover, induce gains of 9% to 20% of SOC. These benefits are higher in clay-rich soils (&gt;30%), due to the increased ability of clay minerals to stabilize organic matter, as well as in sub-humid areas where plant productivity supports organic inputs (<bold>Table 4</bold>).</p>
        <p><bold>Table 4.</bold>Land use transformations and impacts on soil organic carbon (SOC).</p>
        <table-wrap id="tbl4">
          <label>Table 4</label>
          <table>
            <tbody>
              <tr>
                <td>Type of Transformation</td>
                <td>Observed Effects</td>
                <td>Impact on SOC</td>
              </tr>
              <tr>
                <td>Conventional Agriculture and Intensification</td>
                <td>Repeated ploughing; rupture of aggregates; loss of porosity; increased erosion; low organic inputs; decrease in vegetation cover; rapid impoverishment of the upper horizon</td>
                <td>
                  Reduction from 0.3 to 1.2 t∙C∙ha
                  <sup>−</sup>
                  <sup>1</sup>
                  ∙year
                  <sup>−</sup>
                  <sup>1</sup>
                </td>
              </tr>
              <tr>
                <td>Conservation Practices, Agroforestry, and Fallow Land</td>
                <td>Perennial soil cover; continuous biomass inputs; Improved aggregate stability. increased storage in deep horizons; attenuated microclimate; increase in root diversity</td>
                <td>18% - 33% increase in SOC in agroforestry systemsEnhanced deep carbon stability at 30 - 60 cm, especially under shaded cocoa plantations</td>
              </tr>
            </tbody>
          </table>
        </table-wrap>
        <p>The application of biochar and organic amendments is a second major lever. Acidic ferralsols from Cameroon, Ethiopia, and Tanzania show increased stability due to organo-mineral interactions promoted by iron oxides. Agroforestry systems are distinguished by their ability to improve both surface and deep-horizon carbon. The increases are significant: 0.4 to 1.1 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup> at the surface and 0.2 to 0.6 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup> at 30 - 60 cm. The presence of trees improves soil structure, root biomass diversity, and micro-conditions favorable to carbon accumulation. Forest and pasture restoration programs demonstrate a high recovery capacity: 40% to 70% of the initial SOC is restored after 15 to 20 years. This underscores the resilience of tropical ecosystems when anthropogenic pressures decrease.</p>
      </sec>
    </sec>
    <sec id="sec4">
      <title>4. Discussion</title>
      <p>Soil organic carbon (SOC) dynamics in tropical ecosystems in sub-Saharan Africa are the result of a complex interaction between climatic factors, soil intrinsic properties, and anthropogenic pressures. Numerous studies converge to show that climate change plays a structuring role in the mechanisms of carbon decomposition, stabilization, and transfer, but its effects can only be understood by taking into account texture, mineralogy, land uses, and agricultural practices ([<xref ref-type="bibr" rid="B38">38</xref>]; [<xref ref-type="bibr" rid="B7">7</xref>]; [<xref ref-type="bibr" rid="B47">47</xref>]). One of the first major lessons of the corpus analyzed lies in the weight of climatic gradients, in particular precipitation and temperature, which directly modulate biomass production, microbial activity, soil respiration, and mineralization.</p>
      <sec id="sec4dot1">
        <title>4.1. Determining Role of Precipitation and Aridity</title>
        <p>Almost all studies assessing the spatial variability of SOC in tropical areas closely associate organic storage with rainfall levels ([<xref ref-type="bibr" rid="B36">36</xref>]; [<xref ref-type="bibr" rid="B30">30</xref>]; [<xref ref-type="bibr" rid="B39">39</xref>]). The corpus shows that 30 to 55% of the regional variability of the SOC is explained by the precipitation gradient. In semi-arid areas where rainfall is less than 900 mm/year, the water deficit sharply reduces plant biomass, limiting litter inputs and increasing the frequency of fires, which accelerates surface carbon loss ([<xref ref-type="bibr" rid="B5">5</xref>]). Conversely, sub-humid areas, receiving more than 1200 mm/year, show higher stocks, due to more sustained plant productivity, humidity favorable to aggregate formation, and a moderate microclimate ([<xref ref-type="bibr" rid="B16">16</xref>]; [<xref ref-type="bibr" rid="B31">31</xref>]). However, the effect of precipitation is not linear: above a certain threshold, excess moisture induces an acceleration of mineralization or leaching losses, especially in highly weathered ferralsols soils ([<xref ref-type="bibr" rid="B46">46</xref>]; [<xref ref-type="bibr" rid="B10">10</xref>]). Thus, the SOC responds in a non-monotonic way to rainfall variations, according to a water optimum specific to soil-vegetation combinations.</p>
      </sec>
      <sec id="sec4dot2">
        <title>4.2. Thermal Sensitivity as a Major Amplifier of SOC Losses</title>
        <p>The mechanisms linking temperature and SOC are clearly emerging: a 1˚C increase leads to an average loss of 0.21% ± 0.07% of SOC in tropical areas ([<xref ref-type="bibr" rid="B49">49</xref>]; [<xref ref-type="bibr" rid="B1">1</xref>]). Thermal augmentation intensifies heterotrophic respiration, stimulates microbial decomposition, and accelerates the disorganization of organo-mineral complexes ([<xref ref-type="bibr" rid="B17">17</xref>]; [<xref ref-type="bibr" rid="B22">22</xref>]). The work of [<xref ref-type="bibr" rid="B38">38</xref>] indicates that soils dominated by unprotected organic matter, such as Sahelian sands, are particularly vulnerable to temperature increases. Conversely, soils rich in 1:1 clays and iron oxides show higher inertia, as carbon is associated with stable mineral structures ([<xref ref-type="bibr" rid="B40">40</xref>]; [<xref ref-type="bibr" rid="B35">35</xref>]). This increased thermal sensitivity explains the extreme vulnerability of the Sahelian savannah, where the combination of heat + water deficit induces a double stress. Studies by [<xref ref-type="bibr" rid="B15">15</xref>] demonstrate that the increased frequency of heat waves weakens the stability of aggregates, reducing the soil’s ability to encapsulate carbon.</p>
      </sec>
      <sec id="sec4dot3">
        <title>4.3. Mineralogy, Texture, and Organo-Mineral Protection</title>
        <p>Forty-two studies in the corpus emphasize the central role of mineralogical properties in carbon stabilization. Ferralsols soils, which are very common in Central Africa, contain high amounts of iron and aluminum oxides, conferring a significant ability to retain organic compounds via adsorption and co-precipitation ([<xref ref-type="bibr" rid="B20">20</xref>]; [<xref ref-type="bibr" rid="B35">35</xref>]). These soils show carbon stabilization rates 20% to 45% higher than sandy soils under identical climatic conditions. In contrast, sandy soils in the Sahel or southwestern Nigeria lose 30% to 55% of their SOC under prolonged climatic stress, due to their low aggregation capacity and a mineralogy low in protective fine particles ([<xref ref-type="bibr" rid="B30">30</xref>]; [<xref ref-type="bibr" rid="B24">24</xref>]; [<xref ref-type="bibr" rid="B25">25</xref>]). In 41% of the studies, extreme episodes (intense rainfall, severe droughts) induce rapid disintegration of the aggregates, exposing organic matter to accelerated mineralization ([<xref ref-type="bibr" rid="B48">48</xref>]; [<xref ref-type="bibr" rid="B9">9</xref>]). It thus appears that mineral properties determine the degree of amplification or mitigation of climate impacts.</p>
      </sec>
      <sec id="sec4dot4">
        <title>4.4. Anthropogenic Pressures, Conversion, Intensification, and Degradation</title>
        <p>Changes in land use accentuate climate effects. More than 30 studies confirm that forest conversion drastically reduces SOC stocks, with losses ranging from 15 to 45% in rainforests and up to 65% in Cameroonian areas converted to cocoa plantations ([<xref ref-type="bibr" rid="B18">18</xref>]; [<xref ref-type="bibr" rid="B26">26</xref>]). The loss of forest cover reduces litter inputs, disrupts the microclimate, and exposes surface horizons to erosion. Conventional agricultural systems based on frequent ploughing show the highest losses: 0.3 to 1.2 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup>, especially on ferralsols where the fragile structure does not tolerate mechanical disturbance ([<xref ref-type="bibr" rid="B28">28</xref>]; [<xref ref-type="bibr" rid="B45">45</xref>]). The soils of the Plaine du Noun are a good example of these processes, with critical SOC levels often &lt;1% ([<xref ref-type="bibr" rid="B29">29</xref>]; [<xref ref-type="bibr" rid="B23">23</xref>]).</p>
      </sec>
      <sec id="sec4dot5">
        <title>4.5. Sustainable Management Strategies: Effectiveness, Benefits, Limitations</title>
        <p>Work on conservation agriculture shows robust gains, between 9 and 20%, particularly on clay soils in subhumid areas ([<xref ref-type="bibr" rid="B24">24</xref>]; [<xref ref-type="bibr" rid="B27">27</xref>]). According to the work of [<xref ref-type="bibr" rid="B17">17</xref>] and [<xref ref-type="bibr" rid="B12">12</xref>], biochar appears to be one of the most effective long-term interventions with increases of 12% to 60%. The effects are most pronounced in acidic soils where microbial activity is moderate. Agroforestry also strengthens SOC stocks with gains of 0.4 to 1.1 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup> at surface and 0.2 to 0.6 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup> at depth ([<xref ref-type="bibr" rid="B4">4</xref>]; [<xref ref-type="bibr" rid="B31">31</xref>]; [<xref ref-type="bibr" rid="B3">3</xref>]). Restored forest systems show a remarkable ability to recover 40% - 70% of lost carbon in 15 - 20 years ([<xref ref-type="bibr" rid="B16">16</xref>]; [<xref ref-type="bibr" rid="B34">34</xref>]).</p>
      </sec>
      <sec id="sec4dot6">
        <title>4.6. Spatial Heterogeneity and the Contributions of Remote Sensing</title>
        <p>High-resolution mapping reveals a strong regional heterogeneity. Dense moist forests often exceed 60 t∙C∙ha<sup>−</sup><sup>1</sup>, while Sahelian savannahs show values below 15 t∙C∙ha<sup>−</sup><sup>1</sup> ([<xref ref-type="bibr" rid="B8">8</xref>]). Cameroon illustrates an exceptional gradient: &lt;10 t∙C∙ha<sup>−</sup><sup>1</sup> in the northern Sahelian against 80 t∙C∙ha<sup>−</sup><sup>1</sup> in the western mountains ([<xref ref-type="bibr" rid="B44">44</xref>]; [<xref ref-type="bibr" rid="B45">45</xref>]; [<xref ref-type="bibr" rid="B23">23</xref>]). Machine learning-based models achieve accuracies of 62% - 78%, confirming the importance of spectral data and climate variables ([<xref ref-type="bibr" rid="B6">6</xref>]).</p>
      </sec>
    </sec>
    <sec id="sec5">
      <title>5. Conclusion</title>
      <p>This systematic review, based on the analysis of 45 studies published between 2015 and 2025, highlights the decisive influence of climate change on soil organic carbon dynamics in tropical ecosystems in Sub-Saharan Africa. The consolidated results show that 89.4% of the studies establish a direct link between climatic fluctuations and the stability of the SOC, confirming the structural vulnerability of tropical soils to hydroclimatic disturbances. Rainfall gradients explain 30% to 55% of the spatial variability of the SOC, while areas receiving less than 900 mm/year show losses of 0.25% to 0.80% compared to sub-humid areas. Rising temperatures amplify these trends: in 19 quantitative studies, an increase of +1˚C leads to an average reduction of 0.21% ± 0.07% in SOC, reflecting an acceleration of mineralization and CO<sub>2</sub> emission processes.</p>
      <p>The interactions between climate, soil types, and human practices are a major explanatory lever. Sandy or poorly aggregated soils show losses of up to −30% to −55% of SOC under prolonged climatic stress, while soils rich in iron oxides show increased stabilization capacities of 20% to 45%. Land use changes reinforce these mechanisms: deforestation and agricultural conversion reduce stocks by 15% to 65%, while agroforestry and conservation systems allow gains of 18% to 33% and an improvement in deep carbon of 0.2 to 0.6 t∙C∙ha<sup>−</sup><sup>1</sup>∙year<sup>−</sup><sup>1</sup>.</p>
      <p>Thus, the dynamics of the SOC in the region result from a disturbed balance between organic inputs, intensity of mineralization, and mineralogical stabilization capacity. Data show that between 35% and 62% of the net SOC losses observed in tropical landscapes are attributable to climatic pressures, reinforced by unsustainable anthropogenic practices.</p>
      <p>Climate-smart soil-specific management strategies, technologies, innovations, and practices are needed in each SSA country to mitigate the adverse impacts of climate change. However, tropical agricultural soils should be subject to continuous monitoring and assessment to determine which soils and carbon-smart management options should be implemented in a site-specific manner to mitigate soil organic carbon in SSA. This definitely needs a close collaboration between scientists, researchers, policy makers, extension services, and farmers, especially small-scale producers on the ground.</p>
    </sec>
    <sec id="sec6">
      <title>Authors’ Contributions</title>
      <p><bold>Anani Ogou</bold><bold>:</bold> Designed the study, performed the methodology, conducted the review, wrote the original draft of the manuscript, and reviewed and finalized the manuscript. </p>
      <p><bold>Christopher Mubeteneh</bold><bold>Tankou</bold><bold>:</bold> Supervised, performed the methodology, conducted the review, and edited the manuscript.</p>
      <p><bold>Honoré Beyegue-Djonko</bold><bold>:</bold> Performed the methodology and conducted the review.</p>
      <p><bold>Asafor</bold><bold>Henry</bold><bold>Chotangui</bold><bold>:</bold> Performed the methodology and edited the manuscript.</p>
      <p><bold>Martial Georges Ndzana:</bold> Performed the methodology and edited the manuscript.</p>
      <p><bold>Etienne Mboua:</bold> Performed the methodology and edited the manuscript.</p>
      <p><bold>Eric Bertrand Kouam:</bold> Performed the methodology and edited the manuscript.</p>
      <p><bold>Komi Agboka:</bold> Supervised and edited the manuscript.</p>
      <p><bold>Komi Kouma</bold><bold>Mokpokpo</bold><bold>Fiaboe</bold><bold>:</bold> Supervised and edited the manuscript.</p>
    </sec>
    <sec id="sec7">
      <title>Funding</title>
      <p>This article is part of the PhD Thesis activities of Mr. Anani Ogou. It was jointly funded by the European Union (EU) and African Union (AU) throughout the Higher Education Institutions Mobility Program on Climate-Smart Agrifood Systems (CSAS) in East and Central Africa Sub-Regions.</p>
    </sec>
  </body>
  <back>
    <ref-list>
      <title>References</title>
      <ref id="B1">
        <label>1.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Ajayi, A. A., &amp; Okonokhua, B. O. (2024). Spatial Variability of Soil Chemical Properties of an Undulating Site within a University Farm at Okha, near Benin City in Nigeria. <italic>Journal of Applied Sciences and Environmental Management, 28,</italic> 2241-2248. https://doi.org/10.4314/jasem.v28i7.39 <pub-id pub-id-type="doi">10.4314/jasem.v28i7.39</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.4314/jasem.v28i7.39">https://doi.org/10.4314/jasem.v28i7.39</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Ajayi, A.</string-name>
              <string-name>Okonokhua, B.</string-name>
            </person-group>
            <year>2024</year>
            <pub-id pub-id-type="doi">10.4314/jasem.v28i7.39</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B2">
        <label>2.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Akinyemi, F. O., Ghazaryan, G., &amp; Dubovyk, O. (2021). Assessing UN Indicators of Land Degradation Neutrality and Proportion of Degraded Land for Botswana Using Remote Sensing Based National Level Metrics. <italic>Land Degradation &amp; Development, 32,</italic> 158-172. https://doi.org/10.1002/ldr.3695 <pub-id pub-id-type="doi">10.1002/ldr.3695</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1002/ldr.3695">https://doi.org/10.1002/ldr.3695</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Akinyemi, F.</string-name>
              <string-name>Ghazaryan, G.</string-name>
              <string-name>Dubovyk, O.</string-name>
            </person-group>
            <year>2021</year>
            <pub-id pub-id-type="doi">10.1002/ldr.3695</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B3">
        <label>3.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Arthur, A., Acquaye, S., Cheng, W., Dogbatse, J. A., Konlan, S., Domfeh, O. et al. (2022). Soil Carbon Stocks and Main Nutrients under Cocoa Plantations of Different Ages. <italic>Soil Science and Plant Nutrition, 68,</italic> 99-103. https://doi.org/10.1080/00380768.2022.2029219 <pub-id pub-id-type="doi">10.1080/00380768.2022.2029219</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1080/00380768.2022.2029219">https://doi.org/10.1080/00380768.2022.2029219</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Arthur, A.</string-name>
              <string-name>Acquaye, S.</string-name>
              <string-name>Cheng, W.</string-name>
              <string-name>Dogbatse, J.</string-name>
              <string-name>Konlan, S.</string-name>
              <string-name>Domfeh, O.</string-name>
            </person-group>
            <year>2022</year>
            <pub-id pub-id-type="doi">10.1080/00380768.2022.2029219</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B4">
        <label>4.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Asare, R., Asare, R. A., Asante, W. A., Markussen, B. O., &amp; Ræbild, A. (2017). Influences of Shading and Fertilization on On-Farm Yields of Cocoa in Ghana. <italic>Experimental Agriculture, 53,</italic> 416-431. https://doi.org/10.1017/S0014479716000466 <pub-id pub-id-type="doi">10.1017/S0014479716000466</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1017/S0014479716000466">https://doi.org/10.1017/S0014479716000466</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Asare, R.</string-name>
              <string-name>Asare, R.</string-name>
              <string-name>Asante, W.</string-name>
              <string-name>Markussen, B.</string-name>
            </person-group>
            <year>2017</year>
            <pub-id pub-id-type="doi">10.1017/S0014479716000466</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B5">
        <label>5.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Beillouin, D., Corbeels, M., Demenois, J., Berre, D., Boyer, A., Fallot, A. et al. (2023). A Global Meta-Analysis of Soil Organic Carbon in the Anthropocene. <italic>Nature Communi</italic><italic>cations, 14,</italic> Article No. 3700. https://doi.org/10.1038/s41467-023-39338-z <pub-id pub-id-type="doi">10.1038/s41467-023-39338-z</pub-id><pub-id pub-id-type="pmid">37349294</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1038/s41467-023-39338-z">https://doi.org/10.1038/s41467-023-39338-z</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Beillouin, D.</string-name>
              <string-name>Corbeels, M.</string-name>
              <string-name>Demenois, J.</string-name>
              <string-name>Berre, D.</string-name>
              <string-name>Boyer, A.</string-name>
              <string-name>Fallot, A.</string-name>
            </person-group>
            <year>2023</year>
            <elocation-id>No</elocation-id>
            <pub-id pub-id-type="doi">10.1038/s41467-023-39338-z</pub-id>
            <pub-id pub-id-type="pmid">37349294</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B6">
        <label>6.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Bravo-García, J., Camarillo-Naranjo, J. M., Blanco-Velázquez, F. J., &amp; Anaya-Romero, M. (2025). Soil Organic Carbon Mapping through Remote Sensing and in Situ Data with Random Forest by Using Google Earth Engine: A Case Study in Southern Africa. <italic>Land, 14,</italic> Article 1436. https://doi.org/10.3390/land14071436 <pub-id pub-id-type="doi">10.3390/land14071436</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/land14071436">https://doi.org/10.3390/land14071436</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Camarillo-Naranjo, J.</string-name>
              <string-name>Anaya-Romero, M.</string-name>
            </person-group>
            <year>2025</year>
            <elocation-id>1436</elocation-id>
            <pub-id pub-id-type="doi">10.3390/land14071436</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B7">
        <label>7.</label>
        <citation-alternatives>
          <mixed-citation publication-type="web">Chotte, J. L. (2016). <italic>Land Degradation and Climate Change</italic> (p. 45). https://horizon.documentation.ird.fr/exl-doc/pleins_textes/divers16-11/010068505.pdf</mixed-citation>
          <element-citation publication-type="web">
            <person-group person-group-type="author">
              <string-name>Chotte, J.</string-name>
            </person-group>
            <year>2016</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B8">
        <label>8.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Crézé, C., Saatchi, S., Kwon, N., Yang, Y., &amp; Li, S. (2025). <italic>High-Resolution Global Map (100 m) of Soil Organic Carbon Reveals Critical Ecosystems for Carbon Storage</italic> (p. 1-46). Earth System Science Data Discussions.</mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Saatchi, S.</string-name>
              <string-name>Kwon, N.</string-name>
              <string-name>Yang, Y.</string-name>
              <string-name>Li, S.</string-name>
            </person-group>
            <year>2025</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B9">
        <label>9.</label>
        <citation-alternatives>
          <mixed-citation publication-type="web">Deng, X., Shen, J., Lei, Y. B., Sheng, M., Xue, J., &amp; Sun, G. (2025). Revealing Divergence in Soil Carbon Limitation Through Microbial Necromass in Humid and Arid Chronosequences of Alpine Grassland Restoration. SSRN. https://ssrn.com/abstract=5171295</mixed-citation>
          <element-citation publication-type="web">
            <person-group person-group-type="author">
              <string-name>Deng, X.</string-name>
              <string-name>Shen, J.</string-name>
              <string-name>Lei, Y.</string-name>
              <string-name>Sheng, M.</string-name>
              <string-name>Xue, J.</string-name>
              <string-name>Sun, G.</string-name>
            </person-group>
            <year>2025</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B10">
        <label>10.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Desjardins, T., Henry Des Tureaux, T., Mandeng-Yogo, M., &amp; Cetin, F. (2025). Soil Organic Carbon Turnover Following Afforestation of a Savanna Revealed by Particle-Size Fractionation and Natural <sup>13</sup>C Measurements in Ivory Coast. <italic>Land, 14,</italic> Article 535. https://doi.org/10.3390/land14030535 <pub-id pub-id-type="doi">10.3390/land14030535</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/land14030535">https://doi.org/10.3390/land14030535</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Desjardins, T.</string-name>
              <string-name>Tureaux, T.</string-name>
              <string-name>Mandeng-Yogo, M.</string-name>
              <string-name>Cetin, F.</string-name>
            </person-group>
            <year>2025</year>
            <elocation-id>535</elocation-id>
            <pub-id pub-id-type="doi">10.3390/land14030535</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B11">
        <label>11.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Dimobe, K., Zoungrana, B. J.-B., Yoni, M., &amp; Thiombiano, A. (2025). Agroforestry’s Contribution to Sustainable Soil Fertility, Livelihoods, and Carbon Sequestration in Sub-Saharan Africa: A Systematic Review. <italic>International Journal of Agriculture and Biosciences, 14,</italic> 436-446. https://doi.org/10.47278/journal.ijab/2025.013 <pub-id pub-id-type="doi">10.47278/journal.ijab/2025.013</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.47278/journal.ijab/2025.013">https://doi.org/10.47278/journal.ijab/2025.013</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Dimobe, K.</string-name>
              <string-name>Zoungrana, B.</string-name>
              <string-name>Yoni, M.</string-name>
              <string-name>Thiombiano, A.</string-name>
              <string-name>Fertility, L</string-name>
            </person-group>
            <year>2025</year>
            <pub-id pub-id-type="doi">10.47278/journal.ijab/2025.013</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B12">
        <label>12.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Ernest, B., Yanda, P. Z., Hansson, A., &amp; Fridahl, M. (2024). Long-Term Effects of Adding Biochar to Soils on Organic Matter Content, Persistent Carbon Storage, and Moisture Content in Karagwe, Tanzania. <italic>Scientific Reports, 14,</italic> Article No. 30565. https://doi.org/10.1038/s41598-024-83372-w <pub-id pub-id-type="doi">10.1038/s41598-024-83372-w</pub-id><pub-id pub-id-type="pmid">39702623</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1038/s41598-024-83372-w">https://doi.org/10.1038/s41598-024-83372-w</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Ernest, B.</string-name>
              <string-name>Yanda, P.</string-name>
              <string-name>Hansson, A.</string-name>
              <string-name>Fridahl, M.</string-name>
              <string-name>Content, P</string-name>
              <string-name>Karagwe, T</string-name>
            </person-group>
            <year>2024</year>
            <elocation-id>No</elocation-id>
            <pub-id pub-id-type="doi">10.1038/s41598-024-83372-w</pub-id>
            <pub-id pub-id-type="pmid">39702623</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B13">
        <label>13.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">FAO GSP (2021). <italic>Global Soil Organic Carbon Map (</italic><italic>GSOCmap</italic><italic>) v1.5.</italic>FAO.</mixed-citation>
          <element-citation publication-type="other">
            <year>2021</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B14">
        <label>14.</label>
        <citation-alternatives>
          <mixed-citation publication-type="book">FAO GSP (2023). <italic>Soil Organic Carbon Mapping Cookbook</italic>(2nd ed.). FAO.</mixed-citation>
          <element-citation publication-type="book">
            <year>2023</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B15">
        <label>15.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Fonkeng, E. E., Chevallier, T., Sauvadet, M., Enock, S., Rakotondrazafy, N., Chapuis-Lardy, L. et al. (2024). Dynamics of Soil Organic Carbon Pools Following Conversion of Savannah to Cocoa Agroforestry Systems in the Centre Region of Cameroon. <italic>Geoderma</italic><italic>Regional, 36,</italic> e00758. https://doi.org/10.1016/j.geodrs.2024.e00758 <pub-id pub-id-type="doi">10.1016/j.geodrs.2024.e00758</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.geodrs.2024.e00758">https://doi.org/10.1016/j.geodrs.2024.e00758</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Fonkeng, E.</string-name>
              <string-name>Chevallier, T.</string-name>
              <string-name>Sauvadet, M.</string-name>
              <string-name>Enock, S.</string-name>
              <string-name>Rakotondrazafy, N.</string-name>
              <string-name>Chapuis-Lardy, L.</string-name>
            </person-group>
            <year>2024</year>
            <pub-id pub-id-type="doi">10.1016/j.geodrs.2024.e00758</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B16">
        <label>16.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Fujisaki, K., Chevallier, T., Chapuis-Lardy, L., Albrecht, A., Razafimbelo, T., Masse, D., Ndour, Y. B., &amp; Chotte, J.-L. (2018). Soil Carbon Stock Changes in Tropical Croplands Are Mainly Driven by Carbon Inputs: A Synthesis. <italic>Agriculture, Ecosystems &amp; Environment, 259,</italic> 147-158. https://doi.org/10.1016/j.agee.2017.12.008 <pub-id pub-id-type="doi">10.1016/j.agee.2017.12.008</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.agee.2017.12.008">https://doi.org/10.1016/j.agee.2017.12.008</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Fujisaki, K.</string-name>
              <string-name>Chevallier, T.</string-name>
              <string-name>Chapuis-Lardy, L.</string-name>
              <string-name>Albrecht, A.</string-name>
              <string-name>Razafimbelo, T.</string-name>
              <string-name>Masse, D.</string-name>
              <string-name>Ndour, Y.</string-name>
              <string-name>Chotte, J.</string-name>
              <string-name>Agriculture, E</string-name>
            </person-group>
            <year>2018</year>
            <pub-id pub-id-type="doi">10.1016/j.agee.2017.12.008</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B17">
        <label>17.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Gross, A., Bromm, T., &amp; Glaser, B. (2021). Soil Organic Carbon Sequestration after Biochar Application: A Global Meta-Analysis. <italic>Agronomy, 11,</italic> Article 2474. https://doi.org/10.3390/agronomy11122474 <pub-id pub-id-type="doi">10.3390/agronomy11122474</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/agronomy11122474">https://doi.org/10.3390/agronomy11122474</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Gross, A.</string-name>
              <string-name>Bromm, T.</string-name>
              <string-name>Glaser, B.</string-name>
            </person-group>
            <year>2021</year>
            <elocation-id>2474</elocation-id>
            <pub-id pub-id-type="doi">10.3390/agronomy11122474</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B18">
        <label>18.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Ingram, V., Janssen, V., Neh, V. A., &amp; Pratihast, A. K. (2025). Cocoa Driven Deforestation in Cameroon: Practices and Policy. <italic>Forest Policy and Economics, 177,</italic> Article ID: 103533. https://doi.org/10.1016/j.forpol.2025.103533 <pub-id pub-id-type="doi">10.1016/j.forpol.2025.103533</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.forpol.2025.103533">https://doi.org/10.1016/j.forpol.2025.103533</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Ingram, V.</string-name>
              <string-name>Janssen, V.</string-name>
              <string-name>Neh, V.</string-name>
              <string-name>Pratihast, A.</string-name>
            </person-group>
            <year>2025</year>
            <fpage>103533</fpage>
            <elocation-id>ID</elocation-id>
            <pub-id pub-id-type="doi">10.1016/j.forpol.2025.103533</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B19">
        <label>19.</label>
        <citation-alternatives>
          <mixed-citation publication-type="report">IRAD (2021). <italic>Cameroon: SOC Sequestration Potential—National Map &amp; Report.</italic></mixed-citation>
          <element-citation publication-type="report">
            <year>2021</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B20">
        <label>20.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Jiang, Y., Yang, H., Yang, Q., Liu, W., Li, Z., Mao, W. et al. (2021). The Stability of Soil Organic Carbon across Topographies in a Tropical Rainforest. <italic>PeerJ, 9,</italic> e12057. https://doi.org/10.7717/peerj.12057 <pub-id pub-id-type="doi">10.7717/peerj.12057</pub-id><pub-id pub-id-type="pmid">34532159</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.7717/peerj.12057">https://doi.org/10.7717/peerj.12057</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Jiang, Y.</string-name>
              <string-name>Yang, H.</string-name>
              <string-name>Yang, Q.</string-name>
              <string-name>Liu, W.</string-name>
              <string-name>Li, Z.</string-name>
              <string-name>Mao, W.</string-name>
            </person-group>
            <year>2021</year>
            <pub-id pub-id-type="doi">10.7717/peerj.12057</pub-id>
            <pub-id pub-id-type="pmid">34532159</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B21">
        <label>21.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Kebonye, N. M., John, K., Delgado-Baquerizo, M., Zhou, Y., Agyeman, P. C., Seletlo, Z. et al. (2024). Major Overlap in Plant and Soil Organic Carbon Hotspots across Africa. <italic>Science of the Total Environment, 951,</italic> Article ID: 175476. https://doi.org/10.1016/j.scitotenv.2024.175476 <pub-id pub-id-type="doi">10.1016/j.scitotenv.2024.175476</pub-id><pub-id pub-id-type="pmid">39147042</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.scitotenv.2024.175476">https://doi.org/10.1016/j.scitotenv.2024.175476</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Kebonye, N.</string-name>
              <string-name>John, K.</string-name>
              <string-name>Delgado-Baquerizo, M.</string-name>
              <string-name>Zhou, Y.</string-name>
              <string-name>Agyeman, P.</string-name>
              <string-name>Seletlo, Z.</string-name>
            </person-group>
            <year>2024</year>
            <fpage>175476</fpage>
            <elocation-id>ID</elocation-id>
            <pub-id pub-id-type="doi">10.1016/j.scitotenv.2024.175476</pub-id>
            <pub-id pub-id-type="pmid">39147042</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B22">
        <label>22.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Kenfack, F. A. S., Kome, G. K., Ibrahim, A. B., Mandah, V. P., &amp; Bitondo, D. (2024). Soil Organic Carbon Stock Variation under Different Soil Types and Land Uses in the Sub-Humid Noun Plain, Western Cameroon. <italic>Open Journal of Soil Science, 14,</italic> 191-209. https://doi.org/10.4236/ojss.2024.144011 <pub-id pub-id-type="doi">10.4236/ojss.2024.144011</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.4236/ojss.2024.144011">https://doi.org/10.4236/ojss.2024.144011</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Kenfack, F.</string-name>
              <string-name>Kome, G.</string-name>
              <string-name>Ibrahim, A.</string-name>
              <string-name>Mandah, V.</string-name>
              <string-name>Bitondo, D.</string-name>
              <string-name>Plain, W</string-name>
            </person-group>
            <year>2024</year>
            <pub-id pub-id-type="doi">10.4236/ojss.2024.144011</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B23">
        <label>23.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Kome, G. K., Enang, R. K., Yerima, B. P., &amp; Van Ranst, E. (2024). Quantitative Relationships between Munsell Colour Attributes and Organic Carbon in Highly Weathered Tropical Soils. <italic>Geoderma</italic><italic>Regional, 39,</italic> e00898. https://doi.org/10.1016/j.geodrs.2024.e00898 <pub-id pub-id-type="doi">10.1016/j.geodrs.2024.e00898</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.geodrs.2024.e00898">https://doi.org/10.1016/j.geodrs.2024.e00898</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Kome, G.</string-name>
              <string-name>Enang, R.</string-name>
              <string-name>Yerima, B.</string-name>
              <string-name>Ranst, E.</string-name>
            </person-group>
            <year>2024</year>
            <pub-id pub-id-type="doi">10.1016/j.geodrs.2024.e00898</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B24">
        <label>24.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Lekemo, D., Lebeau, T., Amani, I., Kenne, E. R., Tsafack, H. N., Gaudin, P. et al. (2025). Geostatistical and Food Risk Assessment of Soils Contaminated by Trace Elements in the City of Dschang (Cameroon). <italic>Urban Science, 9,</italic> Article 467. https://doi.org/10.3390/urbansci9110467 <pub-id pub-id-type="doi">10.3390/urbansci9110467</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/urbansci9110467">https://doi.org/10.3390/urbansci9110467</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Lekemo, D.</string-name>
              <string-name>Lebeau, T.</string-name>
              <string-name>Amani, I.</string-name>
              <string-name>Kenne, E.</string-name>
              <string-name>Tsafack, H.</string-name>
              <string-name>Gaudin, P.</string-name>
            </person-group>
            <year>2025</year>
            <elocation-id>467</elocation-id>
            <pub-id pub-id-type="doi">10.3390/urbansci9110467</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B25">
        <label>25.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Mandah, V. P., Masso, C., Onana, A. A., Fiaboe, K. K. M., Arthur, E., Giweta, M. et al. (2025a). Soil Organic Carbon and Nutrient Content across Agricultural Systems in the Forest-Savannah Transition Zone of Cameroon. <italic>Soil and Tillage Research, 248,</italic> Article ID: 106458. https://doi.org/10.1016/j.still.2025.106458 <pub-id pub-id-type="doi">10.1016/j.still.2025.106458</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.still.2025.106458">https://doi.org/10.1016/j.still.2025.106458</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Mandah, V.</string-name>
              <string-name>Masso, C.</string-name>
              <string-name>Onana, A.</string-name>
              <string-name>Fiaboe, K.</string-name>
              <string-name>Arthur, E.</string-name>
              <string-name>Giweta, M.</string-name>
            </person-group>
            <year>2025</year>
            <fpage>106458</fpage>
            <elocation-id>ID</elocation-id>
            <pub-id pub-id-type="doi">10.1016/j.still.2025.106458</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B26">
        <label>26.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Mandah, V. P., Tematio, P., Onana, A. A., Fiaboe, K. K. M., Arthur, E., Giweta, M. et al. (2025b). Soil Carbon and Nutrient (NPK) Content in the Tropical Shifting Cultivation System under Indigenous Agricultural Management Practices: A Review. <italic>Environmental Sustainability, 8,</italic> 1-16. https://doi.org/10.1007/s42398-025-00338-y <pub-id pub-id-type="doi">10.1007/s42398-025-00338-y</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1007/s42398-025-00338-y">https://doi.org/10.1007/s42398-025-00338-y</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Mandah, V.</string-name>
              <string-name>Tematio, P.</string-name>
              <string-name>Onana, A.</string-name>
              <string-name>Fiaboe, K.</string-name>
              <string-name>Arthur, E.</string-name>
              <string-name>Giweta, M.</string-name>
            </person-group>
            <pub-id pub-id-type="doi">10.1007/s42398-025-00338-y</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B27">
        <label>27.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Manzeke-Kangara, M. G., Ligowe, I. S., Tibu, A., Gondwe, T. N., Greathead, H. M. R., &amp; Galdos, M. V. (2025). Soil Organic Carbon and Related Properties under Conservation Agriculture and Contrasting Conventional Fields in Northern Malawi. <italic>Frontiers in Soil Science, 4,</italic> Article 1481275. https://doi.org/10.3389/fsoil.2024.1481275 <pub-id pub-id-type="doi">10.3389/fsoil.2024.1481275</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3389/fsoil.2024.1481275">https://doi.org/10.3389/fsoil.2024.1481275</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Manzeke-Kangara, M.</string-name>
              <string-name>Ligowe, I.</string-name>
              <string-name>Tibu, A.</string-name>
              <string-name>Gondwe, T.</string-name>
              <string-name>Greathead, H.</string-name>
              <string-name>Galdos, M.</string-name>
            </person-group>
            <year>2025</year>
            <elocation-id>1481275</elocation-id>
            <pub-id pub-id-type="doi">10.3389/fsoil.2024.1481275</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B28">
        <label>28.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Martinsen, V., Munera-Echeverri, J. L., Obia, A., Cornelissen, G., &amp; Mulder, J. (2019). Significant Build-Up of Soil Organic Carbon under Climate-Smart Conservation Farming in Sub-Saharan Acrisols. <italic>Science of the Total Environment, 660,</italic> 97-104. https://doi.org/10.1016/j.scitotenv.2018.12.452 <pub-id pub-id-type="doi">10.1016/j.scitotenv.2018.12.452</pub-id><pub-id pub-id-type="pmid">30639722</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.scitotenv.2018.12.452">https://doi.org/10.1016/j.scitotenv.2018.12.452</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Martinsen, V.</string-name>
              <string-name>Munera-Echeverri, J.</string-name>
              <string-name>Obia, A.</string-name>
              <string-name>Cornelissen, G.</string-name>
              <string-name>Mulder, J.</string-name>
            </person-group>
            <year>2019</year>
            <pub-id pub-id-type="doi">10.1016/j.scitotenv.2018.12.452</pub-id>
            <pub-id pub-id-type="pmid">30639722</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B29">
        <label>29.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Munjonji, L., Ayisi, K. K., Mudongo, E. I., Mafeo, T. P., Behn, K., Mokoka, M. V. et al. (2020). Disentangling Drought and Grazing Effects on Soil Carbon Stocks and CO <sub>2</sub> Fluxes in a Semi-Arid African Savanna. <italic>Frontiers in Environmental Science, 8,</italic> Article 590665. https://doi.org/10.3389/fenvs.2020.590665 <pub-id pub-id-type="doi">10.3389/fenvs.2020.590665</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3389/fenvs.2020.590665">https://doi.org/10.3389/fenvs.2020.590665</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Munjonji, L.</string-name>
              <string-name>Ayisi, K.</string-name>
              <string-name>Mudongo, E.</string-name>
              <string-name>Mafeo, T.</string-name>
              <string-name>Behn, K.</string-name>
              <string-name>Mokoka, M.</string-name>
            </person-group>
            <year>2020</year>
            <elocation-id>590665</elocation-id>
            <pub-id pub-id-type="doi">10.3389/fenvs.2020.590665</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B30">
        <label>30.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Mustapha, A. A., Abdu, N., Oyinlola, E. Y., &amp; Nuhu, A. A. (2023). Evaluating Different Methods of Organic Carbon Estimation on Nigerian Savannah Soils. <italic>Journal of Soil Science and Plant Nutrition, 23,</italic> 790-800. https://doi.org/10.1007/s42729-022-01082-6 <pub-id pub-id-type="doi">10.1007/s42729-022-01082-6</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1007/s42729-022-01082-6">https://doi.org/10.1007/s42729-022-01082-6</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Mustapha, A.</string-name>
              <string-name>Abdu, N.</string-name>
              <string-name>Oyinlola, E.</string-name>
              <string-name>Nuhu, A.</string-name>
            </person-group>
            <year>2023</year>
            <pub-id pub-id-type="doi">10.1007/s42729-022-01082-6</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B31">
        <label>31.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Nadège, M. T., Louis, Z., Cédric, C. D., Louis-Paul, K. B., Funwi, F. P., Ingrid, T. T. et al. (2019). Carbon Storage Potential of Cacao Agroforestry Systems of Different Age and Management Intensity. <italic>Climate and Development, 11,</italic> 543-554.</mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Louis, Z.</string-name>
              <string-name>Louis-Paul, K.</string-name>
              <string-name>Funwi, F.</string-name>
              <string-name>Ingrid, T.</string-name>
            </person-group>
            <year>2019</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B32">
        <label>32.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Page, M. J., McKenzie, J. E., Bossuyt, P. M., Boutron, I., Hoffmann, T. C., Mulrow, C. D. et al. (2021). Updating Guidance for Reporting Systematic Reviews: Development of the PRISMA 2020 Statement. <italic>Journal of Clinical Epidemiology, 134,</italic> 103-112. https://doi.org/10.1016/j.jclinepi.2021.02.003 <pub-id pub-id-type="doi">10.1016/j.jclinepi.2021.02.003</pub-id><pub-id pub-id-type="pmid">33577987</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.jclinepi.2021.02.003">https://doi.org/10.1016/j.jclinepi.2021.02.003</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Page, M.</string-name>
              <string-name>McKenzie, J.</string-name>
              <string-name>Bossuyt, P.</string-name>
              <string-name>Boutron, I.</string-name>
              <string-name>Hoffmann, T.</string-name>
              <string-name>Mulrow, C.</string-name>
            </person-group>
            <year>2021</year>
            <pub-id pub-id-type="doi">10.1016/j.jclinepi.2021.02.003</pub-id>
            <pub-id pub-id-type="pmid">33577987</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B33">
        <label>33.</label>
        <citation-alternatives>
          <mixed-citation publication-type="web">Python Software Foundation (2025). <italic>Python (Version 3.12.11) [Computer Software].</italic> Python Software Foundation. https://www.python.org/</mixed-citation>
          <element-citation publication-type="web">
            <year>2025</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B34">
        <label>34.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Qasha, V., Manyevere, A., Flynn, T., &amp; Mashamaite, C. V. (2024). Assessing the Impact of Ecological Forest Restoration on Soil Carbon Stocks in Sub-Saharan Africa: A Systematic Review. <italic>Carbon Management, 15,</italic> Article ID: 2404409. https://doi.org/10.1080/17583004.2024.2404409 <pub-id pub-id-type="doi">10.1080/17583004.2024.2404409</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1080/17583004.2024.2404409">https://doi.org/10.1080/17583004.2024.2404409</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Qasha, V.</string-name>
              <string-name>Manyevere, A.</string-name>
              <string-name>Flynn, T.</string-name>
              <string-name>Mashamaite, C.</string-name>
            </person-group>
            <year>2024</year>
            <fpage>240440</fpage>
            <elocation-id>ID</elocation-id>
            <pub-id pub-id-type="doi">10.1080/17583004.2024.2404409</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B35">
        <label>35.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Rabe, M., Droždž, W., Widera, K., Łopatka, A., Lezynski, P., Streimikiene, D., &amp; Bilan, Y. (2022). Assessment of Energy Storage for Energy Strategies Development on a Regional Scale. <italic>Acta Montanistica Slovaca, 27,</italic>163-177.</mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Rabe, M.</string-name>
              <string-name>Widera, K.</string-name>
              <string-name>Lezynski, P.</string-name>
              <string-name>Streimikiene, D.</string-name>
              <string-name>Bilan, Y.</string-name>
            </person-group>
            <year>2022</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B36">
        <label>36.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Reith, J., Ghazaryan, G., Muthoni, F., &amp; Dubovyk, O. (2021). Assessment of Land Degradation in Semiarid Tanzania—Using Multiscale Remote Sensing Datasets to Support Sustainable Development Goal 15.3. <italic>Remote Sensing, 13,</italic> Article 1754. https://doi.org/10.3390/rs13091754 <pub-id pub-id-type="doi">10.3390/rs13091754</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/rs13091754">https://doi.org/10.3390/rs13091754</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Reith, J.</string-name>
              <string-name>Ghazaryan, G.</string-name>
              <string-name>Muthoni, F.</string-name>
              <string-name>Dubovyk, O.</string-name>
            </person-group>
            <year>2021</year>
            <elocation-id>1754</elocation-id>
            <pub-id pub-id-type="doi">10.3390/rs13091754</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B37">
        <label>37.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Rotich, B., Szegi, T., Gelsleichter, Y. A., Fuchs, M., Ocansey, C. M., Phenson, J. N. et al. (2025). Variation in Soil Organic Carbon and Total Nitrogen Stocks across Elevation Gradients and Soil Depths in the Mount Kenya East Forest. <italic>Land, 14,</italic> Article 1217. https://doi.org/10.3390/land14061217 <pub-id pub-id-type="doi">10.3390/land14061217</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/land14061217">https://doi.org/10.3390/land14061217</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Rotich, B.</string-name>
              <string-name>Szegi, T.</string-name>
              <string-name>Gelsleichter, Y.</string-name>
              <string-name>Fuchs, M.</string-name>
              <string-name>Ocansey, C.</string-name>
              <string-name>Phenson, J.</string-name>
            </person-group>
            <year>2025</year>
            <elocation-id>1217</elocation-id>
            <pub-id pub-id-type="doi">10.3390/land14061217</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B38">
        <label>38.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Saiz, G., Bird, M., Wurster, C., Quesada, C. A., Ascough, P., Domingues, T. et al. (2015). The Influence of C <sub>3</sub> and C <sub>4</sub> Vegetation on Soil Organic Matter Dynamics in Contrasting Semi-Natural Tropical Ecosystems. <italic>Biogeosciences</italic><italic>, 12,</italic> 5041-5059. https://doi.org/10.5194/bg-12-5041-2015 <pub-id pub-id-type="doi">10.5194/bg-12-5041-2015</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.5194/bg-12-5041-2015">https://doi.org/10.5194/bg-12-5041-2015</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Saiz, G.</string-name>
              <string-name>Bird, M.</string-name>
              <string-name>Wurster, C.</string-name>
              <string-name>Quesada, C.</string-name>
              <string-name>Ascough, P.</string-name>
              <string-name>Domingues, T.</string-name>
            </person-group>
            <year>2015</year>
            <pub-id pub-id-type="doi">10.5194/bg-12-5041-2015</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B39">
        <label>39.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Sani, R. C. S., Ntoupka, M., Tsozué, D., Vroumsia, T., &amp; Ibrahima, A. (2024). Assessment of Edaphic Conditions in the Mozogo-Gokoro National Park (Sudano-Sahelian Zone of Cameroon). <italic>Environmental and Sustainability Indicators</italic><italic>, 22,</italic> Article 100381. https://doi.org/10.1016/j.indic.2024.100381 <pub-id pub-id-type="doi">10.1016/j.indic.2024.100381</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.indic.2024.100381">https://doi.org/10.1016/j.indic.2024.100381</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Sani, R.</string-name>
              <string-name>Ntoupka, M.</string-name>
              <string-name>Vroumsia, T.</string-name>
              <string-name>Ibrahima, A.</string-name>
            </person-group>
            <year>2024</year>
            <elocation-id>100381</elocation-id>
            <pub-id pub-id-type="doi">10.1016/j.indic.2024.100381</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B40">
        <label>40.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Silatsa, F. B. T., Yemefack, M., Tabi, F. O., Heuvelink, G. B. M., &amp; Leenaars, J. G. B. (2020). Assessing Countrywide Soil Organic Carbon Stock Using Hybrid Machine Learning Modelling and Legacy Soil Data in Cameroon. <italic>Geoderma</italic><italic>, 367,</italic> Article ID: 114260. https://doi.org/10.1016/j.geoderma.2020.114260 <pub-id pub-id-type="doi">10.1016/j.geoderma.2020.114260</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.geoderma.2020.114260">https://doi.org/10.1016/j.geoderma.2020.114260</ext-link></mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Silatsa, F.</string-name>
              <string-name>Yemefack, M.</string-name>
              <string-name>Tabi, F.</string-name>
              <string-name>Heuvelink, G.</string-name>
              <string-name>Leenaars, J.</string-name>
            </person-group>
            <year>2020</year>
            <fpage>114260</fpage>
            <elocation-id>ID</elocation-id>
            <pub-id pub-id-type="doi">10.1016/j.geoderma.2020.114260</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B41">
        <label>41.</label>
        <citation-alternatives>
          <mixed-citation publication-type="web">Spyder Project Contributors (2025). <italic>Spyder IDE (Version 6.0.8) [Computer Software].</italic>Spyder Project. https://www.spyder-ide.org/</mixed-citation>
          <element-citation publication-type="web">
            <year>2025</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B42">
        <label>42.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Taylor, A., Wynants, M., Munishi, L., Kelly, C., Mtei, K., Mkilema, F. et al. (2021). Building Climate Change Adaptation and Resilience through Soil Organic Carbon Restoration in Sub-Saharan Rural Communities: Challenges and Opportunities. <italic>Sustainability, 13,</italic> Article 10966. https://doi.org/10.3390/su131910966 <pub-id pub-id-type="doi">10.3390/su131910966</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.3390/su131910966">https://doi.org/10.3390/su131910966</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Taylor, A.</string-name>
              <string-name>Wynants, M.</string-name>
              <string-name>Munishi, L.</string-name>
              <string-name>Kelly, C.</string-name>
              <string-name>Mtei, K.</string-name>
              <string-name>Mkilema, F.</string-name>
            </person-group>
            <year>2021</year>
            <elocation-id>10966</elocation-id>
            <pub-id pub-id-type="doi">10.3390/su131910966</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B43">
        <label>43.</label>
        <citation-alternatives>
          <mixed-citation publication-type="journal">Tegha, K. C., &amp; Sendze, Y. G. (2016). Soil Organic Carbon Stocks in Mount Cameroon National Park under Different Land Uses. <italic>Journal of Ecology and the Natural Environment, 8,</italic> 20-30.</mixed-citation>
          <element-citation publication-type="journal">
            <person-group person-group-type="author">
              <string-name>Tegha, K.</string-name>
              <string-name>Sendze, Y.</string-name>
            </person-group>
            <year>2016</year>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B44">
        <label>44.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Tsozué, D., Nghonda, J. P., &amp; Mekem, D. L. (2015). Impact of Land Management System on Crop Yields and Soil Fertility in Cameroon. <italic>Solid Earth, 6,</italic> 1087-1101. https://doi.org/10.5194/se-6-1087-2015 <pub-id pub-id-type="doi">10.5194/se-6-1087-2015</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.5194/se-6-1087-2015">https://doi.org/10.5194/se-6-1087-2015</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Nghonda, J.</string-name>
              <string-name>Mekem, D.</string-name>
            </person-group>
            <year>2015</year>
            <pub-id pub-id-type="doi">10.5194/se-6-1087-2015</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B45">
        <label>45.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Tsozué, D., Nghonda, J. P., Tematio, P., &amp; Basga, S. D. (2019). Changes in Soil Properties and Soil Organic Carbon Stocks along an Elevation Gradient at Mount Bambouto, Central Africa. <italic>Catena, 175,</italic> 251-262. https://doi.org/10.1016/j.catena.2018.12.028 <pub-id pub-id-type="doi">10.1016/j.catena.2018.12.028</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1016/j.catena.2018.12.028">https://doi.org/10.1016/j.catena.2018.12.028</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Nghonda, J.</string-name>
              <string-name>Tematio, P.</string-name>
              <string-name>Basga, S.</string-name>
              <string-name>Bambouto, C</string-name>
            </person-group>
            <year>2019</year>
            <pub-id pub-id-type="doi">10.1016/j.catena.2018.12.028</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B46">
        <label>46.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">Tsozué, D., Noubissie, N. M. M., Mamdem, E. L. T., Basga, S. D., &amp; Oyono, D. L. B. (2021). Effects of Environmental Factors and Soil Properties on Soil Organic Carbon Stock in a Natural Dry Tropical Area of Cameroon. <italic>Soil, 7,</italic> 677-691. https://doi.org/10.5194/soil-7-677-2021 <pub-id pub-id-type="doi">10.5194/soil-7-677-2021</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.5194/soil-7-677-2021">https://doi.org/10.5194/soil-7-677-2021</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Noubissie, N.</string-name>
              <string-name>Mamdem, E.</string-name>
              <string-name>Basga, S.</string-name>
              <string-name>Oyono, D.</string-name>
            </person-group>
            <year>2021</year>
            <pub-id pub-id-type="doi">10.5194/soil-7-677-2021</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B47">
        <label>47.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">van Noordwijk, M., Aynekulu, E., Hijbeek, R., Milne, E., Minasny, B., &amp; Dwi Saputra, D. (2023). Soils as Carbon Stores and Sinks: Expectations, Patterns, Processes, and Prospects of Transitions. <italic>Annual Review of Environment and Resources, 48,</italic> 177-205. https://doi.org/10.1146/annurev-environ-112621-083121 <pub-id pub-id-type="doi">10.1146/annurev-environ-112621-083121</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1146/annurev-environ-112621-083121">https://doi.org/10.1146/annurev-environ-112621-083121</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Noordwijk, M.</string-name>
              <string-name>Aynekulu, E.</string-name>
              <string-name>Hijbeek, R.</string-name>
              <string-name>Milne, E.</string-name>
              <string-name>Minasny, B.</string-name>
              <string-name>Saputra, D.</string-name>
              <string-name>Expectations, P</string-name>
            </person-group>
            <year>2023</year>
            <pub-id pub-id-type="doi">10.1146/annurev-environ-112621-083121</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B48">
        <label>48.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">von Fromm, S. F., Doetterl, S., Butler, B. M., Aynekulu, E., Berhe, A. A., Haefele, S. M. et al. (2024). Controls on Timescales of Soil Organic Carbon Persistence across Sub-Saharan Africa. <italic>Global Change Biology, 30,</italic> e17089. https://doi.org/10.1111/gcb.17089 <pub-id pub-id-type="doi">10.1111/gcb.17089</pub-id><pub-id pub-id-type="pmid">38273490</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.1111/gcb.17089">https://doi.org/10.1111/gcb.17089</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Fromm, S.</string-name>
              <string-name>Doetterl, S.</string-name>
              <string-name>Butler, B.</string-name>
              <string-name>Aynekulu, E.</string-name>
              <string-name>Berhe, A.</string-name>
              <string-name>Haefele, S.</string-name>
            </person-group>
            <year>2024</year>
            <pub-id pub-id-type="doi">10.1111/gcb.17089</pub-id>
            <pub-id pub-id-type="pmid">38273490</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
      <ref id="B49">
        <label>49.</label>
        <citation-alternatives>
          <mixed-citation publication-type="other">von Fromm, S. F., Hoyt, A. M., Lange, M., Acquah, G. E., Aynekulu, E., Berhe, A. A. et al. (2021). Continental-Scale Controls on Soil Organic Carbon across Sub-Saharan Africa. <italic>Soil, 7,</italic> 305-332. https://doi.org/10.5194/soil-7-305-2021 <pub-id pub-id-type="doi">10.5194/soil-7-305-2021</pub-id><ext-link ext-link-type="uri" xlink:href="https://doi.org/10.5194/soil-7-305-2021">https://doi.org/10.5194/soil-7-305-2021</ext-link></mixed-citation>
          <element-citation publication-type="other">
            <person-group person-group-type="author">
              <string-name>Fromm, S.</string-name>
              <string-name>Hoyt, A.</string-name>
              <string-name>Lange, M.</string-name>
              <string-name>Acquah, G.</string-name>
              <string-name>Aynekulu, E.</string-name>
              <string-name>Berhe, A.</string-name>
            </person-group>
            <year>2021</year>
            <pub-id pub-id-type="doi">10.5194/soil-7-305-2021</pub-id>
          </element-citation>
        </citation-alternatives>
      </ref>
    </ref-list>
  </back>
</article>