Recently the stratigraphy of the upper Indus basin was presented by [49] [50] . The northern part of the upper Indus show more marine and dolomitic carbonate strata (Figure 3) (Figure 4) while the southern part host dolomitic carbonate along with more arenaceous and evaporitic deposits (Figure 5). The northern part of this belt is represented by Muzaffarabad-Kotli to Nathiagali Abbottabad to Kala Chita to Samana Range to northern Parachinar. The Abbottabad-Nathiagali-Murree section (Figure 4) [15] represents the following lithostratigraphy. Precambrian Hazara Slates consist of mainly slates with some dolomitic limestone and some gypsum. It is correlated with Salkhala slates of Neelam (its type locality is after Salkhala village located on the southern bank of Neelam River) and Reshian valleys and Salt Range Fm in the southern base of Salt Range. Cambrian Abbottabad (dolomite, dolomitic limestone, quartzite/sandstone, with some shale and conglomerate) Formation and Jurassic Datta (sandstone/quartzite) and Samanasuk (limestone and dolomitic limestone with minor shale; dinosaurs trackways horizon is shown in Baroach Nala Malakhel section in Figure 4) Formation are present. Cretaceous Chichali (black to grey shale with glauconitic, chamositic/septechlorite, iron beds with other minerals; well exposed a few kilometers north of Thandiani) and Kawagarh (limestones) formations, and Latest Cretaceous to Earliest Paleocene Indus (laterite, bauxite and fire clay; pisolitic and oolitic nodule common; Cretaceous-Paleogene K-Pg boundary laterite) are present. Paleocene Hangu (coal, shale, sandstone, chamositic ironstone; Hangu Formation is coeval of Patala Formation; shale is common in the Tatta Pani, Kotli

areas) and Lockhart (fine nodular limestone; nodule relatively less than Sakesar/Margala Hill Limestone) are present. Eocene Nammal (shale/muds, marl/limestone), Sakesar (coarse nodular/rubbly limestone; nodule relatively larger than Lockhart Limestone; coeval and also synonym of Margala Hill Limestone), Chorgali (shallow marine green shale and limestone) and Kuldana (alternating red and green shale, limestone and sandstone; transitional marine and continental) formations are present. Miocene-Pliocene Murree Formation (continental red muds, sandstones and some conglomerate); and Quaternary alluvium

including conglomerate, sandstone and mudstone are present. Structurally the road section shows intensive faulting especially thrusts, overturning of strata and some foldings. The major two thrusts are observed in the Abbottabad-Nathiagali section where the Precambrian Hazara Formation is thrusted on the Late Cretaceous Kawagarh and Early Eocene Chorgali Formations respectively. From Nathiagali to Kuldana many thrusts/imbricate faultings are observed in Mesozoic and Cenozoic strata (relatively younger strata). The vertebrate paleontological significance of this area is the findings of the Early Eocene walking whale (mammalian evolution from land to sea) and other mammalian fauna from Kuldana Formation. The general trends of thrust lines are northeast and join with the Jhelum thrust fault in the northeast and also become almost east-west on joining with Kala Chita Range in the southwest. In the Muzaffarabad section (Figure 3) the stratigraphy is almost the same, while no phosphatic localities are discovered from here so far. Here the structure is a Muzaffarabad thrusted anticline. This anticline generally trends northwest to southeast with a doubly plunging nature. The northeastern limb is normal while the southwestern limb is overturned. This anticline is located in the northwestern, northern and northeastern vicinity of Muzaffarabad city. The Kotli section (Figure 3) also represents mostly the same stratigraphy, the only difference is that the thick infratertiary laterite (Indus Formation) and also more thick dolomitic limestone continue from the infratertiary boundary to Cambrian. Apparently, this dolomitic limestone is commonly called Abbottabad dolomitic limestone because no boundaries are found. But due to its thickness than normal, it is possible that the other sequence above the Cambrian may exist here. It needs a detail study to clear whether it is the erosional impact due to tectonic or dolomitic limestone of all or some infra tertiary formations which are present in Nathiagali and surrounding areas. Intense folding and faulting are observed in the Kotli thrusted anticlinal area produced by the convergence of the Indo-Pakistan plate with the Asian plate. Many strike-slip, thrust and imbricate faults are found. The Kotli area consists of many alternations of synclines and anticlines generally trending NNW-SSE. Kotli section (Figure 3) was taken from Tatta Pani anticlinorium. Tatta Pani Anticlinorium represents moderate to high dips, and Cambrian Abbottabad Formation is the core formation. Terminal Cretaceous Indus, Early Paleocene Hangu (Patala), Eocene Nammal, Sakesar, Chorgali and Kuldana and Miocene-Pliocene Murree formations are found on the both limbs. A major thrust fault (after the Jhelum fault) runs from Tatta Pani to Muzaffarabad deduced from previous earthquake intensity, Tatta Pani hot spring and other many features.

The western extension of this northern portion extends upto Koh Safed and Parachinar area. Koh Safed mountain (Figure 4) shows mostly Spinghar Formation (Safed Koh Formation) consisting of possibly Indo-Pakistan shield materials. The Parachinar section (Figure 4) shows stratigraphy representing interfingering of Sulaiman basin and Kohat-Potwar-Kotli basins. The Jurassic Samanasuk limestone (Chiltan Limestone), Cretaceous Chichali shale (Sembar Shale) and Kawagarh Limestone, Early Paleocene Indus Formation laterite and middle to late Paleocene Hangu Formation, Early Eocene Panoba shale (Shaheed Ghat shale), Chashmai and Gurgari Formations (Toi and Kingri Formations), Sheikhan Formation, Bahadurkhel Salt, Jatta Gypsum, Habib Rahi, Domanda, Pirkoh and Drazinda formations, Kuldana Formations, Chinji, Nagri, Dhok Pathan Formations, Lei conglomerate and Soan Formation.

The Salt Range (Katha and Nammal sections; Figure 5), Surghar Range (Figure 5), Khisor Range (Figure 5) and Sheikh Budin hill (Figure 5) sections have mostly the same stratigraphy but some variations (Figure 5). The Baghanwala Formation of Salt Range is replaced by Khisor gypsum in Khisor Range. Further the Hangu Formation of Surghar Range and Nammal sections continue to the eastern Salt Range and Kotli area producing coal, while fine clastic materials and shale increase eastward up to Kotli revealing sand source of Hangu Formation is from south or west and not from east. Its sandstone concentration is being reduced as moving eastward (from Hangu to Katha/Abbottabad and Muzaffarabad/Kotli). But in Salt Range it is commonly called Patala Formation but in actual it is Hangu Formation, however, age may extend up to earliest Eocene. There is no any section that shows the occurrences of both formations in one place. So Hangu name may be used only to avoid any ambiguity in working of Abbottabad coal, Muzaffarabad Coal and also Kotli coal. All these coal belongs to Hangu Formation. Finding of extension of Hangu coal in Tarah area [80] of Khyber agency is encouraging. The stratigraphy of southern part of upper Indus Basin is presented as follows. The oldest Formation is Precambrian Salt Range (evaporitic and terrestrial; pink salt, red shale, marl, gypsum, sandstone). Cambrian Khewra Group represented by Khewra (mainly purple sandstone), Kussak (glauconitic and micaceous sandstone and siltstone interbedded with dolomite), Jutana (mainly dolomite and shale) and Baghanwala (red shale alternated with flaggy sandstone exposed in Salt Range), Khisor (thick gypsum in the base and shale in the upper part exposed in Khisor Range) Formations. Early Permian Nilawahan Group represented by Tobra (freshwater facies of siltstone and shale with pollen and spore flora, and a complex tillites facies of diamictite, sandstone and boulder beds increase in western Salt range and Khisor range), Warcha (speckled sandstone with some shale and coal; Burekhel Permian coal Figure 4); Dandot is local and lateral variation of widely used Warcha, so Dandot is its synonym) and Sardhi (greenish-grey clay with some sandstone, siltstone and limestone) Formations. Middle to Late Permian Zaluch Group represented by Amb (sandstone, limestone and shale), Wargal (mainly limestone and dolomite) and Chidru (shale, quartzose sandstone with minor limestone) Formations, and Triassic Musakhel Group represented by Mianwali (marl, limestone, sandstone, siltstone and dolomite), Tredian (sandstone of terrestrial environments) and Kingriali (dolomite, limestone, dolomitic limestone, marl, sandstone and shale) Formations. Jurassic Surghar Group is represented by Datta (terrestrial deposits like sandstone, shale, ferruginous glass sand and fire clay in the lower part), Shinawari (shale, marl and limestone) and Samanasuk (mainly limestone with minor shale; dinosaur trackways horizon in Surghar Range section, Figure 4 & Figure 5) Formations. Chichali Group represented by Chichali (dark greenish grey glauconitic sandstone and glauconitic shale), Lumshiwal (cross-bedded sandstone and shale of continental origin; coeval to Pab Sandstone of middle and lower Indus basins), Kawagarh (marl and limestone and shale) and Indus (lateritic and bauxitic beds along with pisolitic beds) Formations. The latest Cretaceous or Earliest Tertiary Indus Formation type locality [49] [50] is Narali area (eastern limb) of Tatta Pani Anticlinorium (43K/2), District Kotli, Azad Kashmir (Pakistan). Paleocene Hangu Group is represented by Hangu (sandstone with grey shale and coal) and Lockhart (fine rubbly limestone relative to Sakesar limestone/Margala Hill limestone Formations). Nammal Group is represented by Nammal (shale, marl and limestone) and Sakesar (dominant limestone with subordinate marl; synonym with Margala Hill Limestone) Formations. Nammal and Kuldana Groups are exposed in Potwar-Nathiagali-Muzaffarabad regions while Panoba and Kahan Groups are well exposed in Kohat Region of upper Indus Basin. Panoba and Nammal Groups are correlated with the Chamalang Group of Sulaiman and western Kirthar fold belt and Sohnari Formation of the eastern Kirthar foldbelt. Kuldana Group is correlated with Kahan Group of Kohat sub-basin and Middle Indus basin. Panoba Group represented by Panoba (greenish grey shale with occasional beds of sandstone and limestone), Chashmai (sandstone and green shale; correlated and coeval with Toi Formation of middle and lower Indus basins), Gurguri (sandstone and brown shale; correlated and coeval with Toi Formation of middle and lower Indus basins), Sheikhan (limestone, shale and sandstone; correlated with Drug marl and shale of middle Indus basin), Bahadurkhel Salt (white salt and the upper salt is grey to black) and Jatta Gypsum (gypsum and some clay partings; correlated and coeval with Baska Gypsum). Kahan Group represented by Habib Rahi (mainly light brown limestone with subordinate marl and shale), Domanda (mainly shale with some marl), Pirkoh (mainly white limestone with subordinate marl and shale) and Drazinda (mainly shale with subordinate marl) Formations. Middle to Late Eocene Kuldana Group is represented by Chorgali (green shale and limestone; Kundal coal tar seep Figure 4) and Kuldana (red shale, sandstone and bleached dolomite; show transition toward Murree Formation) Formations. Potwar Group represented by Chinji (red shale and grey to brownish grey sandstone), Nagri (greenish grey sandstone and chocolate brown shale, and conglomerate), Dhok Pathan (monotonous cyclic alternations of grey, greenish grey and brown sandstone, and chocolate color and rusty orange and grey to greenish grey shale, and conglomerate lenses and beds) Formations. Soan Group is represented by Lei Conglomerate (coarse clastics, conglomerate with minor shale and sandstone) and Soan (fine clastics, sandstone, mud and conglomerate) Formations. Subrecent and Recent deposits cover most of the valleys and plain areas. Sheikh Budin hill section (Figure 5) hosts the exposed Tredian, Kingriali, Datta, Shinawari, Samanasuk, Chichali and Lumshiwal Formations along with the Potwar and Soan Group fluvial deposits. Sheik Budin Hill and Khisor Range (form arc shape) and represents a major thrust fault on the southern and eastern end of Range. This thrust joins with Surghar thrust which turns eastward from the northern end of Surghar (west of Kalabagh) and continues up to Nammal, Warcha, Khewra and south of Mirpur and Bhimbar.

The Sulaiman Basin (Middle Indus Basin) consists of about 10 km thick sedimentary sequence (Figure 6) (Figure 7) [49] [50] [21] - [27] . The oldest Sulaiman Group (Permian-Jurassic) represents Permian to Triassic Wulgai (shale with subordinate limestone; Wulgai section (Figure 7), Jurassic Loralai (limestone bed alternating with shale) and Chiltan (mainly limestone) Formations. The Early Cretaceous Parh Group is represented by Sembar (mainly shale with minor marl), Mekhtar (quartzose sandstone; commonly called lower Goru), Goru (shale and marl), and Parh (limestone) Formations. The Late Cretaceous Fort Munro Group is represented by Mughal Kot (Mughalkot, manly shale with subordinate sandstone and rare limestone), Fort Munro (locally developed limestone), Pab (sandstone with subordinate shale) and Vitakri (red muds and greyish white sandstone) Formations. The Mughalkot in Rakhi Gaj section consists of almost mudstone/marl and in Tor Thana (between Mekhtar and Loralai) it consists of large Parh type limestone boulders. The Fort Munro limestone is only found in the Rakhi Gaj (and nearby northern and southern extremity) and Morov (Maximum thick at Dilband-Johan) to Sor Range to Hana Lake (pinched here a few meters) sections. In other places, it is not well recognized. The Sor Member ( [26] , page 529) of Pab Formation is widely exposed from Qila Saifullah to Musafar Pur (Sor Muzghai) (Figure 8) to north of Murgha Kibzai to Manikhawa Shirani to Waziristan [26] [52] . The sandstone of Sor Member hosts the footprints and partial trackways of dinosaurs and pterosaurs (Figure 8). The Kali Member ( [27] , page 903-904; [84] page 5) of Pab Formation is exposed in Tor Thana area (Figure 8), western Sulaiman and also western Kirthar (Figure 9) Foldbelts ( [27] , pages 903-904) where black desert varnish is common and further sandstone matrix is more muddy. The Dhaola Member ( [27] , pages 903-904; [84] , page 5) of Pab Sandstone is found in eastern Sulaiman (Figure 8) and eastern Kirthar (Figure 9) Foldbelts ( [27] , pages 903-904). The sandstone of Dhaola Member hosts the sauropod possibly titanosaurian Dgkhansauroperus footprint (Figure 8). All these three members of Pab Sandstones ( [27] , pages 903-904) are laterally/horizontally variable (Figure 8) and extensions. The Vitakri Formation ( [27] , pages 904, 905) which is deposited just below the Cretaceous-Paleogene boundary (Figure 7) yielded the last dinosaurs, mesoeucrocodiles and pterosaurs, snakes, birds and mammals (see above and below). In the Vitakri and Bara Khadro areas and its vicinity, it is maximum thick, host vertebrate and invertebrate fossils also have conformable lower and upper contacts. Its upper contact shows the Cretaceous-Paleogene boundary (Figure 7). The Paleocene

Figure 7. Stratigraphic Sequence exposed at Rakhi Gaj (Dera Ghazi Khan) (23), Des-Mazar Drik-Sembar-Pazha (Kohlu District) (24), Sanjawi (Ziarat and Loralai) (21), Ziarat Chauter (Ziarat District) (22) and Wulgai Muslimbagh (Qila Saifullah) (18) sections. Not to the scale. For location see Fig. 1. The red dotted line show disconformable contact while black line show conformable contact.

Sangiali Group represents Sangiali (basalticlastic shale and sandstone with Pakiwheel vitakri [64] nautiloids and light brown weathered limestone; local development), Rakhi Gaj (Girdu member, volcaniclastic and hematitic sandstone; Bawata member, alternation of volcaniclastic shale and sandstone), and Dungan (limestone and shale) formations. The Dungan Formation has significant variations from massive limestone (at Harrand, Zinda Pir, Mughalkot and western

Indus suture and its surroundings) to a few thin beds of limestone (at Rakhi Gaj and Mekhtar Spin Ghar section). The Early Eocene.

Chamalang Group represents Shaheed Ghat (shale), Toi (sandstone, shale, rubbly limestone and coal), Kingri (red shale/mud, grey and white sandstone), Drug (rubbly limestone, marl and shale), and Baska (gypsum beds and shale) formations and Early/Middle to Late Eocene Kahan group represents Habib

Rahi (limestone, marl and shale), Domanda (shale with one bed of gypsum), Pir Koh (limestone, marl and shale) and Drazinda (shale with subordinate marl) formations. The Toi and Kingri Formations are found in central, western and northern Sulaiman fold belt and not found in southeastern part of Sulaiman fold belt like Taunsa (Zinda Pir, Pir Gahno and Manjhail areas; Figure 8) to D.G.Khan (Rakhi Gaj; Figure 7) to Rajanpur to Dera Bugti areas. Even these are not found in the southern and eastern parts of Kohlu District like Jhabar, Pinghora, Janthali, Kahan, Tadri, Triman, Makhmar and Mawand areas and also in southern and eastern part of Barkhan District like Lakha Kach Rakhni, Kharcha, Vitakri and Mazara-Dhaola-Andari Range. However, the northern and northwestern part of Kohlu and Barkhan Districts has coal hosting these formations. The Kahan Group yielded the largest marine mammal whales. The Oligocene-Pliocene Vihowa Group represents Chitarwata (grey ferruginous sandstone, conglomerate and mud; host of Buzdartherium large rhino and Asifcroco eucrocodile shown at Zinda Pir Taunsa section, Figure 8), Vihowa (red ferruginous shale/mud, sandstone and conglomerate), Litra ( greenish grey sandstone with subordinate conglomerate and mud), and Chaudhwan (mud, conglomerate and sandstone) formations, and Pleistocene-Holocene Sakhi Sarwar group represents Dada (well-developed conglomerate with subordinate mud and sandstone) and Sakhi Sarwar Formation (poorly developed conglomerate with subordinate mud and sandstone, while in centre of valleys the mud is dominant) concealed at places especially in the valleys and plain areas by the Subrecent and Recent fluvial, eolian and colluvial deposits. The Oligocene terrestrial Chitarwata Formation is only found in the middle Indus basin which yielded the largest land mammals. Foldings (repeated anticlines and synclines) are common in the middle Indus Basin. Anticlinal core represents the oldest rocks as in Shinghar-Takht Sulaiman and Sembar Mazar Drik host the oldest Chiltan Formation (Figure 7). The Gadebar and surroundings anticlines (Loralai region) host the Loralai Formation in the core. The anticlines in the western Indus Suture and eastern vicinity host the Wulgai and Loralai Formations in the core. In Tangi Sar, Andar Pur, Hinglun, Shadiani Fort Munro, Hanki, Tadri Khatan, Triman Khatan, Gamboli anticlines the Early Cretaceous Parh is exposed. In the Pirkoh anticline, the core formation is Habib Rahi exposed in southern limb of Pirkoh anticline. This core formation is becoming younger as shifting anticlines southward and eastward. The South Waziristan section (Figure 6) show Permo-Jurassic strata is almost same or slightly less than southern Sulaiman basin, while Cretaceous and Paleogene strata are pinched and reduced as comparison with Mughalkot Shirani section (Figure 6). The variation from Rakhi Gaj section (Figure 7) to Sanjawi section (Figure 7) to Des-Mazar Drik-Pazha section (Figure 7) to further westward to Ziarat section (Figure 7) and Wulgai section (Figure 7) are presented. Further from north to southward variation can be seen from Toi Nala section (Figure 8) to Zinda Pir section (Figure 8) and Pirkoh-Loti section (Figure 8).

The stratigraphic sequential variations are shown in eastern Kirthar (Sindh Province) and western (Balochistan Province) Kirthar fold belts (Figure 9) [35] [36] [49] [50] .

1) Stratigraphic sequential variations and major anticlinal structures in the lower Indus Basin (western Kirthar Basin; Balochistan Province) of Pakistan.

The western Kirthar foldbelt (western Lower Indus Basin; Balochistan Province) (Figure 9) [35] [36] [49] [50] consists of Permo-Jurassic Sulaiman Group represented by Permo-Triassic Wulgai Formation (shale and minor limestone and marl; shown in Shirinab section (Figure 9). Jurassic Anjira Formation (coeval to Loralai Formation; alternations of limestone and shale) and Chiltan Limestone (Zidi Limestone). The upper limestone bed of Jurassic Chiltan/Zidi Limestone hosts the sauropod Chiltansauroperus footprint (Figure 9). The Early Cretaceous Parh Group is represented by Sembar (mainly shale with minor marl; its lower part hosts Brohisaurus kirthari, Figure 9), Goru (dominantly marl/marly limestone with subordinate shale), and Parh (mainly porcellaneous white limestone) Formations. The Late Cretaceous Fort Munro Group is represented by Mughal Kot (Mughalkot, manly shale with subordinate sandstone and rare limestone), Pab (sandstone with minor shale) and Vitakri (laterite rich in bivalves Pakiring kharzani shells) Formations. The Fort Munro limestone is only found in Morov (Maximum thick at Dilband-Johan section; Figure 9) to Sor Range to Hana Lake (pinched here a few meters) and in the Rakhi Gaj Nala (about 100 m thick) and its nearby northern and southern vicinity. In other places it is not found, only a few limestone beds in upper Mughalkot shales are found in Karkh and Kharzan areas of Khuzdar District. The Pab is exposed only a few meters to 50 m thick in the Karkh area and diminishing toward north (Kharzan) and increasing toward south (Pab Range) and southeast (Bara Laki Range). Further Pab (Kali Member; more muddy sandstone with black desert weathering) exposed in Zidi area in the south of Khuzdar and extends southward to Wad and to Pab Range. Its thickness is increasing southward to Pab Range. Its thickness is maximum at its type locality Pab Range and in the Bara Khadro section of Laki Range. Kali Member [27] is exposed in the western Kirthar foldbelt while Dhaola Member (white quartzose sandstone) [27] is exposed in Eastern Kirthar foldbelt and eastern Sulaiman foldbelt. Its Sor Member is widely exposed from Qila Saifullah to Musafar Pur (Sor Muzghai) to north of Murgha Kibzai to Manikhawa Shirani to Waziristan [26] . These members are laterally variable. The Vitakri Formation which is deposited just below the Cretaceous-Paleogene boundary represented by a thin lateritic bed which host the Pakiring Kharzani. In the Vitakri and Bara Khadro areas and its vicinity, it is maximum thick, host vertebrate and invertebrate fossils also have conformable lower and upper contacts. Its upper contact shows the Cretaceous-Paleogene boundary (Figure 9). The Early Cretaceous shows mostly the same lithological units as Sulaiman basin during Mesozoic and Quaternary but vary in Tertiary strata.

Paleocene Sangiali Group represented by Rakhi Gaj (shale, basaltic beds capped by nautiloid Pakiwheel karkhi Figure 9, sandstone, negligible limestone) and Dungan (mainly limestone with minor shale) Formations. Early Eocene Chamalang Group represented by Shaheed Ghat (shale with minor marl), Toi (sandstone, shale, rubbly limestone and coal) and Kingri (red shale/mud, grey and white sandstone) Formations. The Toi and Kingri Formations are only exposed in northern part of western Kirthar fold belt in the Johan and Dilband (Mastung, Kalat) and surroundings. In the southern part of western Kirthar the only Shaheed Ghat shale is developed and exposed in Kharzan, Karkh and further southward. The middle to Late Eocene Kirthar Group represents Kirthar (limestone, marl and shale) and Gorag (resistant and peak-forming limestone with negligible shale and marl) Formations. The Oligocene Gaj Group represents Nari (sandstone, ironstone, shale, limestone) and Gaj (dominantly shale with rare sandstone, limestone and gypsum) Formations and Miocene-Pliocene Manchar Group represented by Litra (dominant sandstone with subordinate shale and negligible conglomerate) and Chaudhwan (dominant shale with subordinate sandstone and minor conglomerate) Formations which are mainly exposed on the eastern foot of Kirthar Range, while concealed in the plain areas by the Subrecent and Recent fluvial, eolian and colluvial deposits (Figure 9). The variation from Kharzan-Moola to Jhal Magsi section (Figure 9) (western Kirthar foldbelt to Bara Laki section (Figure 9) Karkh section of Khuzdar district show basaltic flows derived from Deccan volcanics at and close to K-Pg boundary (Figure 9) will help to find the age of host and surrounding strata matching with well dated Deccan volcanics.

2) Stratigraphic sequential variations and major anticlinal structures in eastern foldbelt of lower Indus Basin (eastern Kirthar fold belt; Sindh Province) of Pakistan.

The eastern Kirthar fold belt (eastern Lower Indus; Sindh Province) (Figure 9) [35] [36] [49] [50] consists of exposed Fort Munro Group represented by Mughalkot (mainly shale with negligible limestone) and Pab (white Sandstone; its Dhaola Member) and Vitakri Formation (red terrestrial sandstone with subordinate shale) exposed in the core of Laki (Lakhi) anticline and also west of Gorag (commonly called Gorakh a pleasant tourism locality) in Pab Range anticlinorium. The Vitakri Formation is about 50 m thick at Bara Khadro section of Laki Range which is pinching into a thin rusty brown laterite (few centimeters to about estimated 1 meter). This rusty laterite is full of Pakiring kharzani [64] a unique type of bivalves found in Kharzan section (just 3 - 4 km north of Kharzan town). The Bara Khadro stream sand (collected at Tangi at base of Laki Group) yielded red silicified tooth of fish or mammals [38] [51] may be derived from Vitakri Formation due to its red colour. Pab has 3 members namely Dhaola, Kali and Sor Members [27] which are laterally variable. Dhaola Member (white quartzose sandstone) is exposed in Eastern Kirthar foldbelt and eastern Sulaiman foldbelt. Kali Member is exposed in the western Kirthar fold belt and central Sulaiman fold belt. Its Sor Member is widely exposed from Qila Saifullah to Musafar Pur (Sor Muzghai) to north of Murgha Kibzai to Manikhawa Shirani to Waziristan [27] . The Paleocene Ranikot Group represented by Khadro (sandstone, shale, limestone and Deccan volcanics), Bara (sandstone and shale with coal) and Lakhra (limestones and shale) Formations. The Early Eocene Sohnari (lateritic clay and shale, yellow arenaceous limestone pockets, ochre and lignite seams) and middle Eocene Laki (shale, limestone, sandstone, lateritic clay and coal) are found. The Sohnari Formation is widely exposed in the Meting Jhimpir area (southwest of Hyderabad). The Sohnari represents the correlation with its coeval Chamalang Group in the northern Kirthar fold belts and also in Sulaiman fold belt. The Laki Formation correlated with Kirthar Group (Kirthar and Gorag/Gorakh Formations) of western Kirthar and Kahan Group of Sulaiman foldbelt. The Late Eocene-Oligocene Gaj Group represents Late Eocene Nari (sandstone, shale, limestone, ironstone beds) and Gaj (shale with subordinate sandstone and limestone, gypsum bed) formations and Miocene-Pliocene Miocene-Pliocene Manchar Group represented by Litra (dominant sandstone with subordinate shale and negligible conglomerate) and Chaudhwan (dominant shale with subordinate sandstone and minor conglomerate) Formations which are mainly exposed on the eastern foot of Laki Range and also on northern plunges eastern foldbelt (Manchar and its southern vicinity). Pleistocene-Holocene Sakhi Sarwar Group represented by Dada (well developed conglomerate with subordinate mud and sandstone) and Sakhi Sarwar Formation (poorly developed conglomerate with subordinate mud and sandstone, while in centre of valleys, the mud is dominant) rarely exposed in the foot of eastern foldbelt and mostly concealed in plain areas by the Subrecent and Recent (Figure 9). The major variation from eastern Kirthar foldbelt (Bara Laki section; Figure 9) to western Kirthar fold belt is as follows. The stratigraphy and thicknesses of formations of the Bara Laki section and Pab Range section are almost same. While there are many variations from Bara Laki to Gorakh, Karkh and Kharzan Moola sections. The Pab Sandstone is about 400 - 500 m at Bara Laki and Pab Range while it is decreasing upto a few meters in the Karkh and Kharzan areas of Khuzdar district. Further a small thickness of Eocene Sohnari of eastern Kirthar is replaced by more than 500m thick Shaheed Ghat shale and further northward at Johan the Shaheed Ghat, Toi and Kingri formations are developed. In this way Eocene Laki Formation is replaced by a thick sequence of Kirthar and Gorag Formations.

Systematic Paleontology

Dinosauria; Saurischia; Sauropoda; Titanosauriformes; Titanosauria; Poripuchia [58] vide [51] ; Pakisauridae [56] vide [51] ; Isisaurinae [22] ;

Imrankhanhero zilefatmi new genus and new species (Figure 10)

Holotype: Holotype includes a partial right humerus (proximal and mid portion GSP-UM/Sangiali-1124, distal portion GSP/MSM-262-1), distal right femur GSP/MSM-232-1 and proximal right fibula GSP-UM/Sangiali-1117 (Figure 10). The specimen proximal and mid-right humerus GSP-UM/Sangiali-1124 is appropriate in size to pertain to the same specimen as the distal right humerus from the same site. Thus, it is plausible that these two specimens pertain to the same individual. Further this holotypic material is found as surface finds in lower Sangiali with size agreement and no duplication shows belong to a single individual. Holotypic fossil housed in museum of Geological Survey of Pakistan, Quetta, Pakistan except the proximal humerus GSP-UM/Sangiali-1124 and proximal fibula GSP-1117 which are hosted now at the Museum of Paleontology, University of Michigan, USA and will return back to its original position in the museum of Geological Survey of Pakistan, Quetta, Pakistan. Beside this holotypic material a few caudal vertebrae were also found in the same site of lower Sangiali locality but not collected (see below). When the mid Sangiali assemblage was taken, at that time a few bones collected from downstream and upstream which were included in mid Sangiali/eastern Sangiali assemblage. The distance between lower and mid Sangiali sites is about 30 - 50 m. The size of proximal right humerus GSP/Sangiali-1124 and right fibula GSP-UM/Sangiali1117 (may be found at downstream and close to the lower Sangiali site) matches with the size of distal humerus GSP/MSM-262-1 and distal femur GSP/MSM-232-1 collected from lower Sangiali and mid caudal centra observed in lower Sangiali/western Sangiali/north Sangiali site (page 68 of [70] ). The proximal humerus and fibula may be found associated with distal humerus and distal femur. Reference (page 68 of [70] ) mentioned that the Sangiali locality has three main sites from where the dinosaur fossils are found. The western site (lower Sangiali) has associated bones (now holotype of Imrankhanhero zilefatmi) along with slightly tall caudals which are hosted in site. The eastern site (mid Sangiali) produced fragmentary but associated about 100 fossils (now holotype and referred materials of Qaikshaheen masoomniazi). The southern site (upper Sangiali) represents some fossils (vertebrae and limb bones) of stocky limbed Balochisauridae (page 68 of [70] ) which are found as disseminated in the upstream area. Here a humerus may be collected from upstream area of southern Sangiali (nearby to mid Sangiali site) is the holotype (see below) of Ikqaumishan smqureshi.

Type locality, horizon and age: Holotype was found in the lower Sangiali locality (Pakistan dinosaur locality 1; shown in Figure 4 of [51] ) at latitude 29.69812N and longitude 69.39860E of Vitakri dome area, Barkhan district, Balochistan Province, Pakistan. Host horizon of holotype is the upper varicolored but dominantly maroon mud caped by upper sandstone unit of Vitakri Formation (Table 1 of [52] ) of Fort Munro Group [21] [22] [70] . The age of Pab Formation was considered Maastrichtian based on invertebrate such as Orbitoides (Lepidorbitoides) minor ( [88] page 192) and Globotruncana aff. G. linnei, Lituola sp., Omphalocyclusmacropora, Orbitellamedia, Orbitoides sp., and Siderolites sp. [42] . According to stratigraphic position and previously well dated Maastrichtian Pab Formation (lower formation) and well-dated Paleocene formations (upper formations), the age of Vitakri Formation considered the latest Maastrichtian ( [27] pages 904-905, [49] [50] [51] [58] [59] ). The paleogeographic reconstruction indicates that the Kingri coal of Vitakri Formation formed when the area was approximately 9˚S of the equator [89] . Further the Karkh basalt (Figure 9) of Early Paleocene Rakhi Gaj Formation shows its source from major episode of Deccan volcanic eruption. The Vitakri Formation has conformable lower contact with the Pab Formation at Vitakri area reveal the latest Maastrichtian or at least late Maastrichtian. Further the finding of dinosaur fossils conveys at least the Vitakri Formation is not Paleocene and confirmed latest Maastrichtian age. The referred fossils from Zubra peak locality (Figure 1 of [64] ) of Pakistan are found from the latest Maastrichtian Vitakri Formation, and referred tibia K 20/321 (Figures 4a, b, of Plate 1 of [77] ) and right fibula K 27/489 (Figures 5a, b, of Plate 1 of [77] ) from Bara Simla, India [77] reported from latest Maastrichtian Lameta Formation.

Referred specimens: A flattened and transversely compressed almost complete right tibia GSP/MSM-235-7 with anteroposteriorly broad distal tibia, proximal right fibula GSP/MSM-253-7 and right metatarsal GSP/MSM-296-7 are being referred to Imrankhanhero zilefatmi (Figure 10) (Table 4). This referral is based on overlapping fibulae which have the same shape and also appropriate size and features to pertain to the same species. The tibia and metatarsal associated with fibula provide additional informations. These referred material was found from Zubra peak 7 (latitude 29˚43'12"N; longitude 69˚30'16"E) of Dhaola-Gambrak area (shown in Figure 1 of [64] ), Barkhan district, Balochistan, Pakistan. A tibia K 20/321 (Figures 4a, b, of [77] ) and right fibula K 27/489 (Figures 5a, b, of [77] ) from Bara Simla, Jabalpur, India [77] are being referred to Imrankhanhero zilefatmi on shape resembling and size matching with its South Zubra exemplar’s tibia and fibula.

Etymology: Genus name Imrankhanhero honors the Imran Khan who and whose team won the 92 World Cup, hero, for champion and Imran Riaz Khan great journalist and leader of Pakistan to support the poor peoples of Koh Sulaiman Range which are host of fossils. The genus name Imrankhanhero can be pronunciated as Imran Khan Hero. The species name Imrankhanhero zilefatmi honors the Shaheed Zileshah for his sacrifice and Dr. A. N. Fatmi, Former Deputy Director General, Geological Survey of Pakistan, on his contribution regarding Mesozoic paleontology and stratigraphy of Pakistan. The species name zilefatmi can be pronunciated as Zile Fatmi.

Diagnosis: Imrankhanhero zilefatmi medium sized sauropod shares with the Titanosauria on the basis of procoelous caudal centra which are found from this locality. It shares with Poripuchia because of sharing with Pakisauridae. It shares with Pakisauridae on the basis of slender femur (Figure 10) and slightly tall almost squarish and slender caudal. It has following autapomorphies. The humerus of Imrankhanhero zilefatmi is relatively slender, small sized and almost L-shaped. Its humerus has medially inset, sinusoidal and intermediate (between slender and robust) deltopectoral crest and posterior plain surface just below head, while Qaikshaheen masoomniazi has almost laterally set, sinusoidal and robust deltopectoral crest and posterior plain surface just below head, Ikqaumishan smqureshi has medially inset, slender and almost straight deltopectoral crest and posterior plain surface just below head, Gspsaurus pakistani has medially inset, almost straight, slender and a subcircular ridge peak at the base of deltopectoral crest and posterior plain surface just below the head, Pakisaurus balochistani has medially inset, robust and almost straight deltopectoral crest and posterior ridge with lateral and medial triceps, and Balochisaurus malkani has ventrally shifted head and posterior plain surface just below the head. Its distal humerus has no anteriorly expanded radial condyle like Isisaurus colberti while all other titanosaurs from Indo-Pakistan have anteriorly expanded radial condyle. Its distal femoral shaft transverse width is about twice of its anteroposterior width (Figure 10). It has slender femur, while Khanazeem saraikistani, Pakisaurus balochistani and Gspsaurus pakistani have more slender femora, Qaikshaheen masoomniazi has robust femora, Balochisaurus malkani and Marisaurus jeffi have more robust femora. A triangular cavity/hallow is found in the core of femoral distal shaft (Figure 10). Proximal tibia is extremely slender like Khanazeem saraikistani, while Gspsaurus pakistani has robust proximal tibia and Balochisaurus malkani and Qaikshaheen masoomniazi has more robust proximal tibia. Its distal tibia is sub-triangular, anteroposteriorly broad and host centrally situated and wide astragalar fossa (Figure 10) like Khanazeem saraikistani, while Gspsaurus pakistani, Nicksaurus razashahi and Balochisaurus malkani have transversely broad distal tibia with narrow astragalar fossa located in front of small process of distal tibia (Figure 14). Its proximal fibula is more large in anteroposterior width, while Pakisaurus balochistani fibula is large in anteroposterior width, Gspsaurus pakistani and Qaikshaheen masoomniazi fibulae are relatively small in anteroposterior width. For more comparison/diagnosis see below.

Description of elements

Caudal vertebrae: A few small caudal centra found with these holotypic materials are procoelous, slightly tall and subsquarish, with slight ventral reduction (almost no ventral reduction) and neural arch was anteriorly set. Its centra were relatively light and flat laterally than the centra of Qaikshaheen masoomniazi and Ikqaumishan smqureshi and which is heavier and ventrally reduced.

Humerus: A partial right humerus (proximal and mid portion GSP-UM/Sangiali-1124, distal portion GSP/MSM-262-1) was collected (Figure 10). The proximal humerus of Imrankhanhero zilefatmi has L-shaped proximal portion while Isisaurus colberti has relatively flat proximal humerus [65] [90] . The deltopectoral crest of Imrankhanhero zilefatmi is medially inset, intermediate (between robust and slender) and sinusoidal while Qaikshaheen masoomniazi has relatively laterally inset, robust and sinusoidal deltopectoral crest and posterior plain surface just below the head, Ikqaumishan smqureshi has medially inset, slender and straight deltopectoral crest and posterior plain surface just below the head, Gspsaurus pakistani has medially inset, subcircular peak at the base and almost straight deltopectoral crest of proximal humerus, and Pakisaurus balochistani has medially inset and almost straight deltopectoral crest and has posterior ridge (flanked by lateral and medial triceps fossae) just below the head. The shaft section just below the deltopectoral crest shows that mediolateral width is slightly more than anteroposterior depth. The distal humerus show posteriorly well concavity for the adjustment of olecranon process of ulna. The distal humerus of Imrankhanhero zilefatmi has no anteriorly expanded prominent radial condyle like Isisaurus colberti [90] , while Pakisaurus balochistani has anteriorly expanded radial condyle (third condyle other than medial and lateral condyles) located in the mid of transverse distal width of the distal humerus, Balochisaurus malkani has large anteriorly expanded radial condyle which is shifted laterally toward lateral condyle (also flushed with lateral condyle), and Qaikshaheen masoomniazi has relatively small anteriorly expanded radial condyle and migrated toward lateral condyle (but not flushed with lateral condyle). Further six morphs of humeri are mentioned (see below).

Femur: A distal femur GSP/MSM-232-1 (Figure 10) was collected which preserved significant diagnostic features like its slender nature (transverse width slightly more than twice of anteroposterior width) and a triangular cavity/hallow in the core of shaft. Its triangular small cavity or hollow is not found in any other titanosaurs from Indo-Pakistan. Its femoral triangular small cavity or hollow is differentiated from the large hollow cavity of femora of theropods. But here the peripheral bone is twice to thrice thick than triangular core cavity. The cross section of this distal shaft is large sized which shows subrectangular shape. In contrast, the theropods have circular and subcircular hollow and also relatively small cross-sections of femoral bone. Here the femoral difference of theropods and titanosaurs is mentioned because 3 theropods were reported from Vitakri dome area.

Tibia: An almost complete right tibia (GSP/MSM-235-7) (Figure 10, Figure 14) collected from Zubra peak. The proximal part of tibia of Imrankhanhero zilefatmi is flattened (Figure 10, Figure 14) like the proximal flattened tibia of Khanazeem saraikistani (Figure 14) while in contrast the Gspsaurus pakistani has robust proximal tibia (anteroposterior width is slightly to one fourth more than its transverse width) (Figure 14) and Balochisaurus malkani and Qaikshaheen masoomniazi have more robust proximal tibia (where the anteroposterior and transverse widths are subequal) (Figure 14). The distal tibia of Imrankhanhero zilefatmi is subtriangular shaped anteroposteriorly expanded (Figure 10, Figure 14) like the Khanazeem saraikistani (Figure 14), while in contrast the Gspsaurus pakistani, Balochisaurus malkani and Nicksaurus razashahi have transversely expanded and long subparalellogram or suboval shaped distal tibiae (Figure 14). The four morphs (or at least three morphs) of tibiae are found in Pakistan which are very significant for taxonomic comparison. The first morph is the more robust morph represented by subequal transverse and anteroposterior widths (and its long subparalellogram or suboval shaped transversely expanded distal tibia with narrow astragalar scar located in front of short distal process) (Figure 14) with exemplars as Balochisaurus malkani and Qaikshaheen masoomniazi (Figure 11, Figure 14). The second morph is the robust morph represented by anteroposterior width which is round about one and half (1.5) of transverse width, and its long subparalellogram or suboval shaped transversely expanded distal tibia with narrow astragalar scar located in front of short process (Figure 14), with exemplar as Gspsaurus pakistani. The third morph is the slender morph represented by almost flat tibia which have round about twice anteroposterior width than its transverse width and its subtriangular shaped distal tibia has a more transversely expanded width (than Imrankhanhero zilefatmi tibia) and more wide astragalar scar located in the centre of transverse width and also found between the long and short distal processes (Figure 14), with exemplar as Khanazeem saraikistani. The fourth morph is the slender morph represented by almost flat tibia which have round about twice anteroposterior width than its transverse width and its subtriangular shaped distal tibia has relatively less transversely expanded width (than Khanazeem saraikistani tibia) and less wide astragalar scar located in the centre of transverse width and also found between the long and short distal processes (Figure 14), with exemplar as Imrankhanhero zilefatmi. According to reference [85] (Figures 1, 2 of Plate 45 of [85] ) the Saltasaurus juvenile has a more slender tibia than the robust tibia of adult showing ontogenetic growth variations. The author’s observations on plates (Figures 1, 2 of Plate 45 of [85] ) of juvenile and adult tibiae of Saltasaurus which show little variation but both grouped in robust morph (as mentioned above). So no major ontogenetic changes are found in tibiae morphs. The more robust morph of tibiae is also preserved for adult Lohuecotitan pandafilandi [91] from upper Cretaceous of Spain, Lusotitan atalaiensis [92] from late Jurassic of Portugal and Diamantinasaurus matildae [93] [94] from mid-Cretaceous of Australia but unluckily the tibia of Upper Cretaceous juvenile Diamantinasaurus matildae [95] was not recovered. The above 4 morphs reported belong to adult individuals and most of them have almost same size which revealing taxonomic variations (and not ontogenetic growth variation).

Further on observing the cross-section of proximal shafts (Figure 14) of tibiae, there are three prominent morphs of tibiae from Pakistan appeared. The first more robust morph has balloon shaped shaft (Figure 14) with strongly thick posterior portions (and thin anterior portions) with exemplars Qaikshaheen masoomniazi and Balochisaurus malkani (Figure 14). The second robust morph has anticlinal shaped (Figure 14) proximal shaft with central portion thicker than anterior and posterior ends of shaft cross section, with exemplar Gspsaurus pakistani (Figure 14). The third slender morph has oval shaped proximal shaft with subequal thickness at anterior and posterior ends of shaft cross-section (Figure 14) with exemplars Imrankhanhero zilefatmi and Khanazeem saraikistani (Figure 14). All these shafts cross sections were anteroposteriorly long. From Pakistan these 3 morphs reported belong to adult individuals and most of them have almost the same size which revealing taxonomic variations (and not ontogenetic growth variation).

Fibula: The proximal right fibulae GSP-UM/Sangiali-1117 (Figure 10) from lower Sangiali and GSP/MSM-253-7 (Figure 10) from Zubra peak were collected. These proximal fibulae of Imrankhanhero zilefatmi are relatively long/large anteroposterior width than intermediate fibula (between small and long anteroposterior widths) of Pakisaurus balochistani and small fibula (anteroposteriorly expansion is relatively small) of Gspsaurus pakistani and Qaikshaheen masoomniazi. The proximal fibula of Imrankhanhero zilefatmi has relatively smooth lateral and medial sides and has no dorsoventral subparallel fibrous rugosities on the lateral and medial sides (Figure 10), while Pakisaurus balochistani fibula has dorsoventral subparallel fibrous rugosities on the lateral and medial sides. The 3 morphs based on size of anteroposterior expansion of fibulae collected from Pakistan. The first more large morph fibula (Figure 10) (more long anteroposterior expansion) with exemplar is Imrankhanhero zilefatmi. The second large morph fibula (long anteroposterior expansion) with exemplar is

Pakisaurus balochistani. The third small morph fibula (small or less anteroposterior expansion) with exemplars is Qaikshaheen masoomniazi and Gspsaurus pakistani. All these morphs of fibulae have medial diagonal ridge (Figure 10). Here only size of fibula is indicator.

Qaikshaheen masoomniazi has lesser anteroposteriorly width (about 12cm) proximal fibula from mid Sangiali like those of Gspsaurus pakistani has about 13-14cm proximal fibula from Top Kinwa, while Pakisaurus balochistani has larger anteroposteriorly width (about 15cm) proximal fibula from south Kinwa, and Imrankhanhero zilefatmi has more larger anteroposteriorly width (about 18cm) proximal fibula from lower Sangiali and anteroposteriorly width (about 25cm) from Bara Simla. This revealed that the relatively less anteroposterior width bearing proximal fibulae adjusted withmore robust and more transversely expanded proximal tibiae (anteroposterior width is subequal with transverse width), the intermediate anteroposterior width bearing fibulae are adjusted with robust and intermediately transversely expanded proximal tibiae (anteroposterior width is 1.5 of transverse width), and the more larger anteroposterior width bearing fibulae are adjusted with slender and almost flat proximal tibiae (anteroposterior width is twice or more of transverse width). This indicates that the more robust and stronger tibiae are balanced by weak fibulae, the robust and strong tibiae are balanced by intermediate (between weak and strong) fibulae, and the slender tibiae are balanced by strong and robust fibulae to adjust and balance support for body weight.

The shape and size of Chhota Simla fibula resemble with Gspsaurus pakistani Topkinwa left and right fibulae. The references ( [27] , table 3), ( [51] , page 289), ( [52] , page 1045) ( [62] , pages 458-460)and ( [51] , page 422) referred Chhota Simla skeleton to Gspsaurus pakistani on shape resembling of key tibia. But now it is further verified by shape and size resemblance theTopkinwa left and right fibulae with the Chhota Simla left fibula. Now it is plausible that the Chhota Simla limb skeleton is referable to Gspsaurus pakistani. Further the reference ( [96] , page 993) describing the Chhota Simla fibula, claimed in 2011 that only one other fibula [77] has been reported from the Cretaceous of Indo-Pakistan. It seems mistype. While in actual (a pair) left and right proximal fibulaeGSP/MSM-76-16 and GSP/MSM-77-16 were already reported and figured in 2006 (Figure 19b on page 128 of [56] ). Further one author (JAW) also observed these pair of fibulae in 2001 in the museum of Geological Survey of Pakistan, Quetta.

Metatarsal: A partial metatarsal GSP/MSM-296-7 (Figure 10) was collected from Zubra peak area with tibia and fibula within about 2 m length. It is triangular and robust. It has no rugosity which represents it may be a distal metacarpal.

Systematic Paleontology

Dinosauria; Saurischia; Sauropoda; Titanosauriformes; Titanosauria; Poripuchia [58] vide [51] ; Balochisauridae [56] vide [64] ; Balochisaurinae [22] .

Qaikshaheen masoomniazi new genus and new species (Figure 11) (Figure 12).

Holotype: Holotype includes partial posterior cervical centrum GSP-UM/Sangiali-1101, partial cervicodorsal centrum GSP-UM/Sangiali-1176, three partial dorsal vertebrae GSP-UM/Sangiali-1102, GSP-UM/Sangiali-1103 and GSP-UM/Sangiali-1123, dorsal neural arch GSP-UM/Sangiali-1104, three partial caudal vertebrae GSP-UM/Sangiali-1105, GSP-UM/Sangiali-1106, and GSP-UM/Sangiali-1107, distal scapula in 2 pieces GSP-UM/Sangiali-1108 and GSP-UM/Sangiali-1109, and mid scapula GSP-UM/Sangiali-1110; proximal left scapula GSP-UM/Sangiali-1111; left coracoid GSP-UM/Sangiali-1112; proximal right humerus GSP-UM/Sangiali-1113; humerus cross section GSP-UM/Sangiali-1114; distal right humerus GSP-UM/Sangiali-1115; partial ischium GSP-UM/Sangiali-1116; proximal left femur GSP-UM/Sangiali-1118; left distal femur GSP/MSM-1-1; right proximal and mid femur GSP-UM/Sangiali-1119; left proximal tibia GSP-UM/Sangiali-1120; proximal left fibula GSP-UM/Sangiali-1121; mid left fibula GSP-UM/Sangiali-1122 (Figure 11). This assemblage found as surface (10 m * 5 m area) at the single site, size matches (size agreement among holotypic elements) and no duplication among these holotypic elements, show association of a single individual. Fossils are housed in museum of Geological Survey of Pakistan, Quetta, Pakistan but now present at the Museum of Paleontology, University of Michigan, USA and will return back to its original position in the museum of Geological Survey of Pakistan, Quetta, Pakistan. Reference (page 68 of [70] ) mentioned the collection from 3 sites of Sangiali Locality 1 (see above).

Type locality, horizon and age: Holotype was found in the mid Sangiali locality (Pakistan dinosaur locality 1) with an area of about 10 m long and 5 m wide area bounded by latitudes 29.69810N and 29.69812N and longitudes 69.39872E and 69.39882E (shown in Figure 4 of [51] ) of Vitakri dome area, Barkhan district, Balochistan Province, Pakistan. This type locality is found in the northern part of Vitakri dome. Host horizon of holotype is the upper varicolored but dominantly maroon mud caped by upper sandstone unit of Vitakri Formation (Table 1 of [52] ) of Fort Munro Group [21] [22] [70] . According to stratigraphic position and previously well-dated Maastrichtian Pab Formation (lower formation) and well-dated Paleocene formations (upper formations), the age of Vitakri Formation considered the latest Maastrichtian ( [27] pages 904-905, [49] [50] [51] [58] [89] ). For detail please see above. The referred fossil assemblage from mid Bor locality (Figure 4 of [51] ) of Pakistan is found from the latest Maastrichtian Vitakri Formation, and referred right humerus from Bara Simla locality, Jabalpur, India (Figures 2a, b, of Plate 5 of [77] ) and referred right humerus from Rahioli locality, Gujarat state, India (GSI 20012, Figure 4 of Plate 1 of [78] ) and referred femur GSI/WR/M-90-84 from Rahioli, western India [65] [78] [79] are reported from the latest Maastrichtian Lameta Formation.

Referred specimens: The referred materials from mid Bor locality (shown in Figure 4 of [51] ) consists of cervical vertebra GSP/MSM-359-2, cervicodorsal vertebra GSP/MSM-120-2, distal part of cervical rib GSP/MSM-187-2, dorsal vertebrae GSP/MSM-121-2, distal thoracic rib GSP/MSM-301-2, GSP/MSM-122-2, GSP/MSM-123-2, GSP/MSM-124-2, GSP/MSM-125-2, GSP/MSM-441-2, a pair of sacral vertebrae GSP/MSM-135-2, caudal vertebrae GSP/MSM-41-2 and GSP/MSM-42-2, GSP/MSM-360-2, GSP/MSM-302-2, neural spine GSP/MSM-792-2, distal rib/neural spine GSP/MSM-784-2, prezygapophyses and postzygapophyses GSP/MSM-560-2; parts of sternal GSP/MSM-565-2 and GSP/MSM-1004-2, a coracoid GSP/MSM-560-2, humerus parts GSP/MSM-559-2, GSP/MSM-287-2, GSP/MSM-363-2, GSP/MSM-362-2; left and right ulnae GSP/MSM-573-2 and GSP/MSM-271-2, distal ulna GSP/MSM-852-2; proximal metacarpals GSP/MSM-295-2, GSP/MSM-279-2, GSP/MSM-685-2, GSP/MSM-566-2, GSP/MSM-278-2, GSP/MSM-686-2, GSP/MSM-1029-2, GSP/MSM-688-2; distal metacarpals GSP/MSM-277-2, GSP/MSM-1028-2, GSP/MSM-285-2, GSP/MSM-370-2, GSP/MSM-684-2, GSP/MSM-687-2, GSP/MSM-361-2, GSP/MSM-683-2, proximal ischium GSP/MSM-184-2, a femur (proximal half GSP/MSM-178-2 and distal half GSP/MSM-182-2), and metatarsals GSP/MSM-643-2, GSP/MSM-1031-2 and GSP/MSM-1030-2 (Figure 12). These fossils are housed in the Museum of Geological Survey of Pakistan, Quetta, Pakistan. The mid Bor Pakistani assemblage is referred to Qaikshaheen masoomniazi due to overlapping typical robust femur and vertebrae with appropriate size and shape resembling. The mid Bor assemblage belongs to plausibly a single individual and a single species based on size agreement and finding within about 10*3m area. A femur GSI/WR/M-90-84 from Rahioli, western India [65] [78] [79] is being referred to Qaikshaheen masoomniazi due to shape resembling and appropriate size. A right humerus from Bara Simla, Jabalpur, India (Figures 2a, b, of Plate V of [77] and a humerus from Rahioli, Gujarat state, India (GSI 20012, Figure 4 of Plate 1 of [78] ) are being referred to Qaikshaheen masoomniazi due to overlapping humeri and also shape resembling and appropriate size.

Etymology: Genus name Qaikshaheen honors the Qaed AllamaIqbal and Imran Khan, abbreviated as Qaik (QAIK), shaheen, Urdu/Saraiki for king. The genus name Qaikshaheen can be pronunciated as Qaik, Shaheen. Here honored are Allama Iqbal great leader of Pakistan, and both Imran Khan Former Prime Minister of Pakistan and Imran Riaz Khan great journalist and leader to support poor peoples of Sulaiman fold belt which host these fossils. The species name Qaikshaheen masoomniazi is after the masoom, Urdu/Saraiki for innocent, niazi, after the honorable Former Prime Minister of Pakistan who supported the poor peoples of Koh Sulaiman area (host of bones). The species name masoomniazi can be pronunciated as Masoom-Niazi.

Diagnosis: Qaikshaheen masoomniazi medium sized sauropod shares with the Titanosauria as procoelous caudals and anteriorly oriented neural arch. It shares with Poripuchia (all tail have procoelous centra) due to finding of procoelous distalmost caudals besides anterior, mid and posterior procoelous tail vertebrae. Qaikshaheen masoomniazi diagnosed as below. Qaikshaheen masoomniazi has proximal scapula (including glenoid and adjoining part) with almost straight alignment and trend with shaft, while Pakisaurus balochistani has laterally or outwardly deflected proximal scapula. Its coracoid has no lip while Pakisaurus balochistani coracoid has lip (Figure 13). Its proximal humerus has almost laterally inset robust deltopectoral crest (Figure 11) which is not found in any titanosaurs especially from Indo-Pakistan (comparison see in description). It has robust femur (Figure 11) (Figure 12) while Balochisaurus has more robust femur, and Imrankhanhero zilefatmi, Gspsaurus pakistani, Pakisaurus balochistani and Khanazeem saraikistani have more slender femora. Its femoral head is slightly inclined dorsally like Gspsaurus pakistani, Nicksaurus razashahi, Balochisaurus malkani and Qaikshaheen masoomniazi, while Pakisaurus balochistani and Khanazeem saraikistani have strongly inflected and inclined upward head. Its proximal femur has a straight profile of lateral deflection like Gspsaurus pakistani, Balochisaurus malkani and Nicksaurus razashahi, while Khanazeem saraikistani has lateral concavity between the head and lateral bulge of proximal femur. Its proximal tibia is more robust like Balochisaurus malkani, while Gspsaurus pakistani has robust and Khanazeem saraikistani and Imrankhanhero zilefatmi have slender and flat tibiae. For detail comparison of tibia and other elements, see below.

Description of elements

Vertebrae: The cervicals are opisthotic, broad transversely and ventral smooth surface like the cervicals of Gspsaurus pakistani. Its cervicals are relatively smaller than the Gspsaurus pakistani cervicals. Cervical rib is broad V-shaped. Its dorsals are opisthotic, slightly transversely broad and have smooth surface in anterior dorsal and has ventral ridge in posterior dorsal. It is believed that the dorsal with smooth ventral surface is found in the anterior and may be in mid, while ventral keel ridge is found in the posterior dorsal deduced from sacral vertebrae which have ventral keel. Its sacral vertebrae are broad, pneumatic and have ventral keel. Its caudals are procoelous, short and have significant ventral reductions (Figure 11) (Figure 12). Its distalmost caudals (Figure 12) are procoelous and grouped in Poripuchian titanosaurs. Lithostrotian did not have procoelous distalmost caudals while Poripuchian had all tail procoelous and more or most derived titanosaurs. Further the reference ( [97] , page 19) mentioned in 2019 “Regardless of the taxonomic validity of the named Pakistani species a considerable amount of titanosaur materials has been collected from the Pab Formation by the Geological Survey of Pakistan. Of the elements that have been published and the unpublished elements from Pakistan, that we have studied first hand, there are no anterior caudal vertebra and only one anterior dorsal vertebra that is directly comparable to the Indian elements described here” ( [97] , page 19). Their claim [97] for no any anterior caudal reported from Pakistan is mistyped. While in actual anterior caudal vertebra GSP/MSM-43-15 ofBalochisaurus malkani (Figures 9-12 of [56] and GSP/MSM-11-4 of Pakisaurus balochistani (Figures 5-8 of [56] and GSP/MSM-23-3 and GSP/MSM-24-15 of Sulaimanisaurus gingerichiwere already reported and figured in 2006 [56] and GSP/MSM-219-19 and MSM-218-19 of Maojandino alami [61] and GSP/MSM-347-4n and GSP/MSM-348-4n of Nicksaurus razashahi [61] . Further their claim that only one anterior dorsal vertebra from Pakistan is again mistyped. Some dorsal vertebrae along with cervical and sacral vertebrae were also reported from Pakistan in 2006 [87] .

Scapula: Qaikshaheen masoomniazi has proximal scapula (Figure 11) with almost straight alignment with mid shaft like Nicksaurus razashahi and Gspsaurus pakistani, while Pakisaurus balochistani has laterally or outward deflected proximal scapula from mid-shaft. Its distal scapular blade is plate-like like while mid slightly concaved.

Coracoid: The glenoid of coracoid of Qaikshaheen masoomniazi has no lip (Figure 11), while Pakisaurus balochistani coracoid has lip (Figure 13).

Humerus: Its proximal humerus (Figure 11) has medially inset (relatively close to lateral end) of ventral end of deltopectoral crest, while laterally set and flushed with lateral humerus of mid deltopectoral crest and turned slightly to medial its proximal portion, as a whole resulting sinusoidal/wavy nature. These features are nor known in other Titanosaurs from Indo-Pakistan subcontinent. Its deltopectoral crest is robust. The Qaikshaheen masoomniazi has almost laterally inset, robust and sinusoidal deltopectoral crest and posteriorly plain surface (Figure 11), while Ikqaumishan smqureshi has medially inset, slender and straight deltopectoral crest and posterior plain surface just below the head and prominent medial process of proximal humerus, Gspsaurus pakistani has medially inset, a ridge in the base and straight deltopectoral crest, Isisaurus colberti has medially inset deltopectoral crest and Pakisaurus balochistani from Pakistan and Diamantinasaurus matildae from Australia [93] [94] [95] and Rapetosaurus krausei from Madagascar [98] has medially inset and straight deltopectoral crest and posterior mid ridge and triangular shape of shaft cross section just below the deltopectoral crest of proximal humerus. Its distal humerus has expanded radial condyle but shape and position is different than Pakisaurus balochistani and Balochisaurus malkani (Figure 2 of [52] ). Its radial condyle is relatively small and positioned just close to lateral small condyle (Figure 11), while Pakisaurus balochistani radial condyle is positioned almost in the centre of distal transversal profile (Figure 2 of [52] ) and Balochisaurus has relatively large radial condyle positioned just close to lateral small condyle (Figure 2 of [52] ). The sixth morph of humeri are presented as below.

Ulna: A pair of proximal ulnae GSP/MSM-573-2 and GSP/MSM-271-2 (Figure 12) collected from mid Bor locality assemblage. The proximal ulna is rugose and bears a prominent olecranon process. The proximal ulna represents a triradiate structure. The ulna is gracile. There is a marked concavity on the proximal lateral side for the adjustment of radius. It has also concavity on the medial side and a relatively small concavity on the posterior side. The relative length of ulnar proximal condylar processes is unequal. Many ulnae were collected from Pakistan but provide no taxonomic major differentiations.

Metacarpals: Many metacarpals were collected from mid Bor (Figure 12). Metacarpals are elongated and robust. Distal and proximal condyles are not divided.

Ischium: Partial ischium GSP/MSM-184-2 (Figure 12) is found from mid Bor locality. The proximal part is thin and platy having iliac condyle, and long and thin acetabular glenoid and then articular surface for pubis. It is only first ischium found from Pakistan. This ischium is remarkably slender than the ischium of Isisaurus colberti [90] . This ischium has low pubis and iliac peduncle. Further theanteroposterior width of pubis and iliac peduncle are almost subeaqual.Its anteroposteriorly width of iliac peduncle is 12 cm, acetabular glenoid is 21cm and pubic peduncle is 12 cm. Its maximum ateroposterior width at the base of iliac and pubic peduncle is about 30 cm. This maximum thickness is ventrally being reduced. It became anteroposterior 18 cm, just 12 cm below the ventral border of iliac and pubic peduncle.Its acetabular glenoid is concave shaped. It is a generally fan shape. Its anteroposterior width of pubic attachment is small or narrow (12 cm), while the Rapetosaurus krauseihas relatively long anteroposterior width [98] .

Femur: Its femora are robust (Figure 11) (Figure 12) while Balochisaurus malkani has more robust femur, and Pakisaurus balochistani, Khanazeem saraikistani, Imrankhanhero zilefatmi and Gspsaurus pakistani have more slender femora. Its femoral head is slightly inclined like Gspsaurus pakistani, Balochisaurus malkani and Nicksaurus razashahi while Pakisaurus balochistani and Khanazeem saraikistani has strongly inflected and inclined upward head. Its dorsolateral corner is straight like Gspsaurus pakistani, Balochisaurus malkani and Nicksaurus razashahi, while Khanazeem saraikistani has a concavity between the posterolateral corner and lateral buldge. For comparison the three morphs of femora (based on slenderness/robustness) found from Pakistan are being presented here. The more robust morph of femur is represented by transverse width considerably less than twice of anteroposteriorly width, with exemplars from Pakistan are Balochisaurus malkani (and may be Marisaurus jeffi and Nicksaurus razashahi). The robust morph of femur is represented by transverse width slightly less than twice of anteroposteriorly width, with exemplars from Pakistan are Qaikshaheen masoomniazi. The slender morph of femur is represented by transverse width equal or more than twice of anteroposteriorly width, with exemplars from Pakistan are Gspsaurus pakistani, Pakisaurus balochistani, Khanazeem saraikistani and possibly Imrankhanhero zilefatmi.

Tibia: The preserved left proximal tibia (GSP-UM/Sangiali-1120) of Qaikshaheen masoomniazi shows more robust morph and its cross section in the proximal shaft show balloon shaped with posterior portion strongly more thick than anterior end of cross section of shaft (Figure 11, Figure 14). The base of its cnemial crest is relatively thick. The tibia of Qaikshaheen masoomniazi (Figure 11, Figure 14) is more robust and heavy like the tibia of Balochisaurus malkani,

while Gspsaurus pakistani has robust (Figure 14) and Khanazeem saraikistani and Imrankhanhero zilefatmi have slender flat tibiae (Figure 14). Its four morphs are described above. Its proximal tibia is extremely transversely expanded approaching to its subequal transverse and anteroposterior widths (Figure 14) like Balochisaurus malkani (Figure 14) from latest Cretaceous of Pakistan, Lohuecotitan pandafilandi [91] from upper Cretaceous of Spain, Lusotitan atalaiensis [92] from late Jurassic of Portugal, and Diamantinasaurus matildae

[93] [94] from mid Cretaceous of Australia; while Gspsaurus pakistani has intermediately expanded proximal tibia with anteroposterior width is round about one and half (1.5) of transverse width (Figure 14), and Khanazeem saraikistani and Imrankhanhero zilefatmi have flat proximal tibia with strongly twice anteroposterior width than transverse width (Figure 14).

Fibula: Its fibula (Figure 11) has relatively small anteroposterior width like the fibulae of Gspsaurus pakistani, while Pakisaurus balochistani has relatively intermediate width (between small and long/larger anteroposterior widths) of proximal fibula, and Imrankhanhero zilefatmi has long/larger anteroposterior width of proximal fibula. For detail see above.

Metatarsals: Many metatarsals were collected from mid Bor (Figure 12). Metatarsals are elongated, broad and have rugose articular surfaces. Distal condyle is wide, have rugosities and shape is slightly divided and has central concavity

Figure 14. Row 1, p1,2, proximal right tibiae (proximal with upper shaft) GSP/MSM-72-2 from lower Bor of Khanazeem saraikistani in dorsal and lateroventral views; p3,4, proximal right tibiae (proximal with upper shaft) GSP/MSM-73-16 from Topkinwa of Gspsaurus pakistani in dorsal and lateroventral views; p5,6, proximal left tibiae (proximal with upper shaft) GSP/MSM-246-15 of Balochisaurus malkani from south Mari Bohri and GSP-UM/Sangiali-1120 of Qaikshaheen masoomniazi from mid-Sangiali in ventral views. Row 2, proximal left tibia with upper shaft GSP/MSM-181-2 of Gspsaurus pakistani from east Bor in dorsal and lateral views. Row 3, p1,2, anteroposteriorly broad subtriangular shaped distal tibiae GSP/MSM-186-2 of Khanazeem saraikistani from mid Bor and GSP/MSM-235-7 of Imrankhanhero zilefatmi from Zubra peak in lateral views; p3, transversely long subparalellogram or suboval shaped right distal tibia GSP/MSM-710-19 of Gspsaurus pakistani from Alam in ventral view; p4,5, transversely long subparalellogram or suboval shaped right distal tibia GSP/MSM-75-9 of Balochisaurus malkani from Grut in ventral and laterodorsal views. Scale, each black or white/yellow digit is 1 cm for row 1,3. Scale is in inches for row 2.

to adjust the large oval ungual for attachment or relevant phalange. A proximal and distal end is expanded with triangle shape. Metatarsals represent irregular concave proximal and distal expanded surface.

Systematic Paleontology

Dinosauria; Saurischia; Sauropoda; Titanosauriformes; Titanosauria; Poripuchia [58] vide [51] ; Gspsauridae [61] vide [51] ; Gspsaurinae [61] vide [51] ;

Ikqaumishan smqureshi new genus and new species (Figure 13).

Holotype: A diagnostic partial humerus (GSP-UM/Sangiali-1125) (Figure 13) is designated as holotype. This holotypic humerus found as 3 pieces but fit into an almost complete proximal and mid humerus GSP-UM/Sangiali-1125 (Figure 13), so belong to a single individual. Fossil is housed in museum of Geological Survey of Pakistan, Quetta, Pakistan but now present at the Museum of Paleontology, University of Michigan, USA and will return back to its original position in the museum of Geological Survey of Pakistan, Quetta, Pakistan. Further from upper Sangiali (south Sangiali) locality, a few ventrally reduced caudal vertebrae, and limb fossils (page 68 of [70] ) were found in Vitakri Formation. After the collection of about 2800 bones, the bone assemblages and many bones were observed and left in site for future study, but not collected during later field visits. These observations are mentioned in 2009 by reference (page 68 of [70] ) and in many other reports. This is also discussed (see above). During collection of mid sangiali assemblage, a few bones were found and collected in the vicinity of mid Sangiali. Holotypic humerus of Ikqaumishan smqureshi seems to be taken from upstream area (just above the mid Sangiali). In this way a holotypic proximal humerus of Ikqaumishan smqureshi was considered from South Sangiali/upper Sangiali. Further among mid Sangiali collections, a few bones were in duplication and their size will not match. One simple armor plate (GSP/MSM-8-1; GSP-BC-001/01) from southern bank of Sangiali stream of titanosaur (and two crocodilian specimens GSP/MSM-9-1 (GSP-BC-002/01) and GSP/MSM-10-3 (GSP-BC-003/01) from Shalghara, see below) were received by Dr David Krauss, Bostan College, USA (during his field visit in 2001) from Geological Survey of Pakistan for preparation ( [66] , pages 532-534; [99] , page 245). This simple armor plate GSP/MSM-8-1 (GSP-BC-001/01) found from southern bank of Sangiali stream of Sangiali dinosaur locality, may belong to a same and single individual of holotypic humerus (see detail below).

Type locality, horizon and age: Holotype was found in the upper Sangiali locality (Pakistan dinosaur locality 1; shown in Figure 4 of [51] ) at latitude 29.69809N and longitude 69.39882E of Vitakri dome area, Barkhan district, Balochistan Province, Pakistan. This holotype is found just above/south of the mid Sangiali locality shown in Figure 4 of [51] , the area just above/south of mid Sangiali in mapped triangle (Figure 4 of [51] ) is included in south Sangiali/upper Sangiali. Host horizon of holotype is the upper varicolored but dominantly maroon mud caped by upper sandstone unit of Vitakri Formation (Table 1 of [52] ) of Fort Munro Group [21] [22] [70] . According to stratigraphic position and previously well dated Maastrichtian Pab Formation (lower formation) and well dated Paleocene formations (upper formations), the age of Vitakri Formation considered latest Maastrichtian ( [27] pages 904-905, [49] [50] [51] [58] [89] ). For detail please see above. Further referred right and left humeri (78 cm length) GSI type no 20010 (Figure 1(a), Figure 1(b) of Plate 1 of [78] ) and GSI 20011 [78] from Rahioli locality of Gujarat state, India reported from the latest Maastrichtian Lameta Formation.

Referred specimens: A proximal right humerus GSP/MSM-237-10 from Rahi Wali locality (Figure 1 of [64] ) of Mazara-Dhaola-Andari Range of Pakistan is being referred to Ikqaumishan smqureshi based on shape resembling and posterior plain surface just below the head. Further right and left humeri (78 cm length) GSI type no 20010 (Figure 1(a), Figure 1(b) of Plate 1 of [78] ) and GSI 20011 [78] reported from Rahioli locality, Gujarat state, India from Lameta Formation are being referred to Ikqaumishan smqureshi based on shape resembling and also matching features medially inset, slender and almost straight deltopectoral crest and posterior smooth surface just below head. The referred humeri from India provide additional informations especially about distal humeri.

Etymology: The genus name Ikqaumishan honors the Former Prime Minister Imran Khan and great journalist Imran Khan (both abbreviated as I.K.) to support poor peoples of Sulaiman Basin of Pakistan which is the host of these fossils, and qaumi, Urdu/Saraiki for nation, and shan, Urdu/Saraiki for respect. The genus name Ikqaumishan can be pronunciated as I.K.-Qaumi-Shan. The species name Ikqaumishan smqureshi honors the honorable Shah Mahmood (Qureshi) and Saeed Murad (Murad Saeed) (abbreviated as sm, for both) formers federal ministers of Pakistan, qureshi for tribe name of Mohtram Shah Mahmood Qureshi for his support to the Saraiki, Balochi and other poor peoples of Koh Sulaiman Range and Sulaiman foldbelt of Pakistan (host of fossils). The species name Ikqaumishan smqureshi can be pronunciated as S.M. Qureshi.

Diagnosis: Ikqaumishan smqureshi medium sized sauropod shares with the Titanosauria on the basis of procoelous caudal vertebrae which are found from this upper Sangiali (south Sangiali) locality. It shares with Poripuchia because of sharing with Gspsauridae. It shares with Gspsauridae on the basis of medially inset and almost straight deltopectoral crest. Its autapomorphy shown as; Ikqaumishan smqureshi has medially inset, slender and almost straight deltopectoral crest and strong medial expansion of medial process of proximal humerus (Figure 13), while this prominent medially expanded medial process is not found in any Indo-Pakistani titanosaurs. Its deltopectoral crest is more slender (and straight) than the slender deltopectoral crest of Gspsaurus pakistani, intermediate (between slender and robust) and sinusoidal deltopectoral crest of Imrankhanhero zilefatmi (and may be Isisaurus colberti) and robust deltopectoral crest of Qaikshaheen masoomniazi and Pakisaurus balochistani. Its deltopectoral crest is relatively thin at the base/ventral portion than the deltopectoral crest of Gspsaurus pakistani specimen NHMUK R5931 from Chhota Simla ( [51] , page 289; [52] , page 1045; ( [27] , see Table 3; [62] , page 459) which are more thick and sub-circular shaped peak forming. Its humerus has posteriorly plain surface while Pakisaurus balochistani has posterior ridge just below the head with triceps fossae. Further diagnosis and comparison are described as below.

Description of elements

Caudal vertebrae: A few caudal vertebrae found in the south Sangiali (upper Sangiali) are procoelous, and show ventral reduction. Its caudal centrum is heavy like those of Balochisaurids and Gspsaurids, while Pakisaurids have relatively light and ventrally not reduced. Its caudal centrum is ventrally reduced and more heavy than those of Imrankhanhero zilefatmi.

Humerus: A holotypic partial right humerus (proximal and mid portion GSP-UM/Sangiali-1125) (Figure 13) collected from upper Sangiali (possibly just above the mid Sangiali). A partial humerus GSP/MSM-237-10 from Rahi Wali (Figure 13) is referred to Ikqaumishan smqureshi on the shape resembling and matching of posterior plain surface just below the head and shaft cross section just below the deltopectoral crest. This referred humerus is providing more significant features which portion is not preserved in holotype. Further right and left humeri (78cm length) GSI type no 20010 (Figure 1(a), Figure 1(b) of Plate 1 of [78] ) and GSI 20011 [78] reported from Rahioli locality, Gujarat state, India are being referred to Ikqaumishan smqureshi based on matching features medially inset, slender, relatively thin and almost straight deltopectoral crest and also posterior smooth surface just below the head.

Ikqaumishan smqureshi has relatively slender and distinguished deltopectoral crest and expansion of medial process of proximal humerus which are not shown in any Indo-Pakistani titanosaurs (as shown above). Ikqaumishan smqureshi has medially inset, slender and almost straight deltopectoral crest and medially prominent expanded medial process of proximal humerus (Figure 13) and posterior plain surface just below the head (Figure 13), while Pakisaurus balochistani has medially inset, robust and almost straight deltopectoral crest and posterior dorsoventral ridge (with left and right triceps fossa) starting just below the head and runs downward to mid shaft, Gspsaurus pakistani has medially inset and almost straight deltopectoral crest with subcircular ridge at base and relatively less medial expansion of proximal humerus, Qaikshaheen masoomniazi has almost laterally inset, robust and sinusoidal deltopectoral crest, Balochisaurus malkani has smooth posterior plain just below the head and relative more downward migrated head, Imrankhanhero zilefatmi has medially inset, intermediate (between slender and robust) and sinusoidal deltopectoral crest, and Isisaurus colberti has medially inset and relatively robust deltopectoral crest. The Rahiwali humerus (Figure 13) of Ikqaumishan smqureshi has prominent medial expansion than other humeri of Qaikshaheen masoomniazi, Imrankhanhero zilefatmi and Pakisaurus balochistani, Gspsaurus pakistani and Isisaurus colberti. The shaft section just below the deltopectoral crest shows that mediolateral width is slightly more than anteroposterior depth. The distal humerus show posterior concavity for the adjustment of olecranon process of ulna and anteriorly expanded radial condyle known from Rahioli referred humeri. The posterior incline thin ridges on holotypic humerus of Ikqaumishan smqureshi has no medial and lateral triceps fossae and may be due to diagenetic and or tectonic compression.

Six morphs of humeri from Indo-Pakistan appeared on comparison of the humerus of Ikqaumishan smqureshi with other humeri of different genera. The first morph is represented by medially inset deltopectoral crest and posterior ridge just below the head and continue to midshaft flanked by lateral triceps fossa and medial triceps fossa with exemplar as Pakisaurus balochistani (row 2 of Figure 13) from Pakistan and also from India [52] [78] and adult and juvenile Diamantinasaurus matildae from Australia [93] [94] [95] and Rapetosaurus krausei from Madagascar [98] . Triceps fossa is site for attachment of tridirectional movement muscles. The finding of this morph of humeri from Indo-Pakistan (South Asia) and also from Australia and Madagascar show their land associations. The second morph is represented by medially inset slender deltopectoral crest and posterior smooth plain surface just below the head, and prominent medial expansion of medial process of proximal humerus (Figure 13) with exemplar Ikqaumishan smqureshi. The third morph is represented by medially inset, intermediate (between slender and robust) deltopectoral crest and posterior smooth plain surface just below the head (Figure 13) with exemplar Imrankhanhero zilefatmi. The fourth morph is represented by medially inset, subcircular ridge at the ventral base and straight deltopectoral crest and posterior smooth plain surface just below the head, and relatively less (than Ikqaumishan smqureshi humerus) medial expansion of medial process of proximal humerus with exemplar Gspsaurus pakistani from Indo-Pakistan subcontinent, and Atsinganosaurus velauciensis from France [99] . The fifth morph is represented by relatively laterally set robust deltopectoral crest and posterior smooth plain surface just below the head (Figure 11) with exemplar Qaikshaheen masoomniazi. The sixth morph is represented by downward migrated head of proximal humerus, and plain surface just below the head and anteriorly expanded relatively large radial condyle flushed with lateral condyle (Figure 7, row 5 of [64] ; Figure 7, rows 1, 2 of [63] ) with exemplar Balochisaurus malkani. Further variation in distal humeri was also discussed (see above).

Limb bones: Limb bones along with ventrally reduced caudal vertebrae [70] were found in the upper Sangiali (south Sangiali) suggesting their assignment to stocky titanosaurian sauropods.

Osteoderm: One simple armor plate or osteoderm GSP/MSM-8-1 (GSP-BC-001/01) of titanosaur (found from southern bank of Sangiali stream) was received by Dr David Krauss, Bostan College, USA (during his field visit in 2001) from Geological Survey of Pakistan for preparation ( [66] , pages 532-534), along with the following crocodile fossils (see below). This armor plate of titanosaur was collected from the southern Bank of Sangiali stream. Further this armor plate site located in the south of mid Sangiali assemblage. The catchment area of this armor plate which is found on the southern bank of stream represents southern Sangiali. So it is found very close to holotypic humerus GSP-UM/Sangiali-1125 of Ikqaumishan smqureshi. So it is possible it may be belong to single individual of the holotypic humerus. This armor bone is thin and subcircular and without internal duct. Ikqaumishan smqureshi armor bone is simple, small and thin subcircular plate without internal duct, while Gspsaurus pakistani has slightly more thick and subrectangular mosaic armor plates (GSP/MSM-83-16 and GSP/MSM-1035-16) (Figure 5 of [62] ) from Topkinwa which bears many irregular internal ducts, Gspsaurus pakistani has large, thick and heavy ellipsoidal plate (GSP/MSM-85-4) with median groove found from south Kinwa (Figure 7 of [62] ), Nicksaurus razashahi has large, thick and heavy ellipsoidal plate without median groove/cut (GSP/MSM-84-7) found from south Zubra (Figure 9 of [63] ), and Balochisaurus malkani has the mosaic type small sub oval armour bone (GSP/MSM-1095-17) (Figure 9 of [63] ) from Dada Pahi 17 locality. The large and heavy ellipsoidal osteodermal plate (GSP/MSM-84-7) of Nicksaurus razashahi resemble with Malawisaurus [100] of Malawi, Africa. The large and heavy ellipsoidal plate (GSP/MSM-85-4) of Gspsaurus pakistani resemble with titanosaur (plate 77 of [85] ) from Argentina, South America. These large heavy ellipsoidal osteoderms show paleobiogeographic link with Africa and South America of Gondwana. In this way majority of bones show affinity to Gondwana and while some also show affinity to Laurasia.

Paleobiogeographic Implications

Here first I described the useful feedback on fossils of Shalghara locality and their paleobiogeographic affinity and then other significant key elements of titanosaurian sauropods from Pakistan with paleobiogeographic implications. Only one crocodile Mithasaraikistan ikniazi [51] and one theropod Shansaraiki insafi [52] were named from Shalghara locality 3 (Pakistan dinosaur locality no. 3) with Latitude 29.68288 and Longitude 69.38008 (N 29˚40'58"; E 69˚22'48") (as shown in Figure 4 of [51] ) of Vitakri dome area, Barkhan district, Balochistan. Mithasaraikistan ikniazi [51] based on Snout GSP/MSM-4-3 and dentary ramus GSP/MSM-139-3. Further a partial humerus GSP/MSM-9-3 (GSP-BC-002/01) and three articulated vertebrae GSP/MSM-10-3 (GSP-BC-003/01) of crocodiles (see below) found from Shalghara locality are being referred to Mithasaraikistan ikniazi on the basis of finding from one Shalghara locality, more holotypic elements (than single referred dentary symphysis of Pabwehshi Pakistanensis from Shalghara [67] ) and also size agreement. Mithasaraikistan ikniazi [51] is diagnosed on the strongly medially inset of premaxillary teeth, while Pabwehshi pakistanensis has relatively laterally flushed premaxillary teeth. One simple armor plate GSP/MSM-8-1 (GSP-BC-001/01) of titanosaur found from southern bank of Sangiali stream, a partial humerus GSP/MSM-9-3 (GSP-BC-002/01) and three articulated vertebrae GSP/MSM-10-3 (GSP-BC-003/01) of crocodiles found from Shalghara locality were received by Dr David Krauss, Bostan College, USA (during his field visit in 2001) from Geological Survey of Pakistan for preparation in 2001 ( [66] , pages 532-534; [101] , page 245).

Shansaraiki insafi was based on specimens GSP/MSM-140-3 (symphysis), GSP/MSM-5-3 (mid ramus with partial teeth bases) and GSP/MSM-57-3 (dorsal vertebra) that were found from the Shalghara locality 3 with Latitude 29.68288 and Longitude 69.38008 (N 29˚40'58"; E 69˚22'48") (as shown in Figure 4 of [52] ) of Vitakri dome area, Barkhan district, Balochistan. The number following the locality such as Shalghara 3 or Shalghara locality 3 and last number of most of specimen numbers represents the locality number of Pakistan Dinosaur Localities PDL-1 to PDL-25. The following two feedbacks and positive criticism were provided regarding Shansaraiki. The Prehistoric Wiki entered the Shansaraiki insafi on web is encouraging, but mentioned that the Shansaraiki “named from GSP/MSM-140-3 (symphysis), GSP/MSM-5-3 (mid ramus with partial teeth bases) and GSP/MSM-57-3 (dorsal vertebra) that were found from various areas within the Vitakri dome. Doubt about these remains connectiveness and validity has been raised, since they were found apart and are extremely fragmentary” (Shansaraiki in prehistoric.wiki.fandom.com). On designing the type locality of Shansaraiki ( [52] , page 1067), it is clearly mentioned the Shalghara locality 3 of Vitakri Dome area with one site grid coordinates. They may be confused from Shalghara locality 3, here 3 represents Pakistan Dinosaur Locality 3 (among PDL-1 to PDL-25; pages 66 and 68 of [52] ). It is also cleared in caption of Figure 2 of ( [52] , page 1046). Further Shalghara locality has only one site from where the fossils were collected and reported. It is no doubt these fossils were found surface find fragmentary from only one site of Shalghara Locality but not from different areas of Vitakri dome. Vitakri dome has more than 10 kilometer area (5*2km area; see in Figure 2 of [70] ) which hosts many localities and also many type localities and many sites within a locality at places. So it is expected the Prehistoric Wiki will correct this fact to avoid any ambiguity. The Tyler Greenfield on twitter questioned regarding Shansaraiki holotype and diagnosis “Why are a dentary symphysis and dorsal vertebrae that were not found together referred to the same taxon? Why were they named at all when they have no diagnostic features? (December 5, 2022 on twitter by @Tyler Greenfield). The reply of his first question is found in Holotype title ( [52] , page 1067). Holotypic bones found as fragmentary on the surface of same Formation, same locality, same time, fit size and no duplication shows associated bones of single individual ( [52] , page 1067). He may also be confused from Shalghara locality 3, here 3 represents Pakistan Dinosaur Locality 3. It is cleared in caption of Figure 2 of ( [52] , page 1046). Further it is necessary to mention that the dentary symphysis and dorsal vertebrae are not found together in animal but they were located apart and have some distance between them. The reply of his second question is found in diagnosis and comparison ( [52] , pages 1069-1072). From Indo-Pakistan this theropod is unique (see diagnosis and comparison in pages 1069-1072 of [52] ). The dentary ramus of Shansaraiki insafi does not bear lateral step while Rajasaurus narmadensis [102] represent lateral step on dentary ramus. Further its dentary and its anterior weak symphysis matches closely with dentary of Australovenator wintonensis from Australia [93] . The suggestions regarding third question is to go through in detail study of more recent research on Pakistani dinosaur taxa (3 theropods and 13 titanosaurian sauropods), although proliferated but most of these are diagnostic. These both feedbacks are very positive and useful for our understandings of Pakistani dinosaurs.

From Pakistan many diverse and significant fossils for taxonomy and paleobiogeographic implications were found from the latest Maastrichtian Vitakri Formation. The nature of V-shaped teeth arcade of Gspsaurus pakistani and 40⁰ anterior slope of cranial face of Saraikimasoom vitakri poripuchian titanosaur are endemic in Titanosaurs especially in derived titanosaurs. The procoelous tail vertebrae are common while the Balochisaurus malkani and Marisaurus jeffi have first biconvex caudals like those of Pellegrinisaurus and Neuquensaurus from Argentina, South America, and Alamosaurus from USA, North America [62] [63] [103] . This feature revealed the Paleobiogeographic link of Indo-Pakistan subcontinent with both Gondwana (southern lands) and Laurasia (northern lands). This feature and element shows worldwide distribution. The Khanazeem saraikistani has unique and endemic slender femur with inflected head and a lateral concavity between the proximolateral corner of head and lateral bulge.

The Pakisaurus balochistani is diagnostic on two major key elements and their features as inflected head of femur and posterior ridge (Figure 13) just below the head of humerus along with lateral and media triceps fossae [27] [52] . This humeral key element of Pakisaurus balochistani Pakistani titanosaur ( [27] and references therein) closely resemble with mid Cretaceous and late Cretaceous Australian titanosaur Diamantinasaurus matildae [94] [95] and late Cretaceous Malagasy (Madagascar) Rapetosaurus krausei [98] .

The most robust tibiae of Balochisaurus malkani ( [27] and references therein) and Qaikshaheen masoomniazi (Figure 11) shows extreme transverse expansion resulting subequal transverse and anteroposterior widths of proximal tibia (Figure 14), resemble closely with Lohuecotitan pandafilandi [91] from upper Cretaceous of Spain (Europe), Lusotitan atalaiensis [92] from late Jurassic of Portugal (Europe), Diamantinasaurus matildae [93] [94] from mid Cretaceous of Australia and Janenschia robusta [104] from the late Jurassic of Tanzania (Africa).

The robust tibiae of Gspsaurus pakistani has intermediately transverse expanded proximal tibia with anteroposterior width which is 1.5 of transverse width like those of Atsinganosaurus velauciensis titanosaur from the upper Cretaceous of southern France [99] and Mendozasaurus neguyelap titanosaur from the upper Cretaceous of Mendoza Province, Argentina [105] and Saltasaurus loricatus from late Cretaceous of Argentina (plate 45 of [85] ). This feature and elements show both Gondwana and Laurasian worldwide distribution.

The slender and almost flat tibiae of Imrankhanhero zilefatmi (Figure 11) and Khanazeem saraikistani closely resemble with Igai semkhu [106] from the late Cretaceous of Egypt. Further the distal tibiae of Imrankhanhero zilefatmi (Figure 11) and Khanazeem saraikistani is anteroposteriorly expanded and subtriangular (Figure 11) (Figure 14), like those of Igai semkhu [106] from the late Cretaceous of Egypt, while Gspsaurus pakistani, Nicksaurus razashahi and Balochisaurus malkani have transversely expanded and broad distal tibia (Figure 14). This reveals that Imrankhanhero zilefatmi and Khanazeem saraikistani of Indo-Pakistan subcontinent have close relation with Igai semkhu [106] of Egypt (Africa).

Most of the key elements of Indo-Pakistani titanosaurs resemble more closely with the Gondwanan lands than those of the Laurasian lands. Briefly this can be interpreted that during early Cretaceous Indo-Pakistan subcontinent was associated with Australia, Madagascar, Africa and also South America. Later on all these lands of Gondwana separated and travelled toward their own present destinations. This separation during Early Cretaceous resulted genetic seed dispersal and changes occurred during migration.