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T. J. Givnish, K. J. Sytsma, J. F. Smith and W. J. Hann, “Thorn-Like Prickles and Heterophylly in Cyanea: Adaptations to Extinct Avian Browsers on Hawaii?” Proceedings of National Academy of Sciences of the USA, Vol. 91, No. 7, 1994, pp. 2810-2814. doi:10.1073/pnas.91.7.2810

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