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T. Sang, D. J. Crawford and T. F. Stuessy, “Documentation of Reticulate Evolution in Peonies (Paeonia) Using Internal Transcribed Spacer Sequences of Nuclear Ribosomal DNA: Implications for Biogeography and Concerted Evolution,” Proceedings of the National Academic of Sciences of the United States of America, Vol. 92, No. 15, 1995, pp. 6813-6817. doi:10.1073/pnas.92.15.6813
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