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Collares-Pereira, M.J. and Madeira, J.M. (1990) Cytotaxonomical Studies in Iberian Cyprinids. III. Karyology of Portuguese Populations of Barbus Cuvier, 1817, with Some Reconsiderations on the Karyological Evolution of Cyprinidae. Caryologia, 43, 17-26.
https://doi.org/10.1080/00087114.1990.10796983

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