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Génier, F. and Moretto, P. (2017) Digitonthophagus Blathasar, 1959: Taxonomy, Systematics, and Morphological Phylogeny of the Genus Revealing an African Species Complex (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa, 4248, 1-110.
https://doi.org/10.11646/zootaxa.4248.1.1

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