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Kotiaho, J.S. and Simmons, L.W. (2003) Longevity Cost of Reproduction for Males but No Longevity Cost of Mating or Courtship for Females in the Male-Dimorphic Dung Beetle Onthophagusbinodis. Journal of Insect Physiology, 49, 817-822.
http://dx.doi.org/10.1016/S0022-1910(03)00117-3

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