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Tyler, P.A., Bronsdon, S.K., Young, C.M. and Rice, A.L. (1995) Ecology and Gametogenic Biology of the Genus Umbellula (Pennatulacea) in the North Atlantic Ocean. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 80, 187-199.
http://dx.doi.org/10.1002/iroh.19950800207

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