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Wordsworth, B.P., Lanchbury, J.S., Sakkas, L.I., Welsh, K.I., Panayi, G.S. and Bell, J.I. (1989) HLA-DR4 Subtype Frequencies in Rheumatoid Arthritis Indicate That DRB1 Is the Major Susceptibility Locus within the HLA Class II Region. Proceedings of the National Academy of Sciences of the United States of America, 86, 10049-10053.
http://dx.doi.org/10.1073/pnas.86.24.10049

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