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J. H. Graham, K. Shimizu, J. M. Emlen, D. C. Freeman and J. Merkel, “Growth Models and the Expected Distribution of Fluctuating Asymmetry,” Biological Journal of the Linnean Society, Vol. 80, No. 1, 2003, pp. 57-65.
http://dx.doi.org/10.1046/j.1095-8312.2003.00220.x

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