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S. D. Allison, “Cheaters, Diffusion and Nutrients Constrain Decomposition by Microbial Enzymes in Spatially Structured Environments,” Ecology Letters, Vol. 8, No. 6, 2005, pp. 626-635.
http://dx.doi.org/10.1111/j.1461-0248.2005.00756.x

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